Salix rotundifolia |
Salix orestera |
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least willow, round-leaf willow |
gray-leaf Sierra willow, gray-leafed Sierra willow, Sierra willow |
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Habit | Plants 0.005–0.05 m, (dwarf), forming clones by rhizomes. | Plants 0.5–2 m. Stems: branches dark red-brown or yellow-brown, not to strongly glaucous, glabrous; branchlets yellow-brown or red-brown, (not or weakly glaucous), pilose or pubescent, hairs straight, wavy, or geniculate, (inner membranaceous bud-scale layer free, separating from outer layer). | ||||
Stems | erect; branches yellow-green, yellow-brown, or gray-brown, glabrous; branchlets yellow-brown or red-brown, glabrous; branches and branchlets sometimes weakly glaucous. |
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Leaves | (marcescent but not skeletonized), stipules usually absent or rudimentary, rarely present on late ones; petiole (convex, or shallowly to deeply grooved, flat), 0.4–4.6(–5.5) mm, (glabrous adaxially); largest medial blade (2 pairs of secondary veins arising at or close to base, arcing toward apex) broadly elliptic, subcircular, or circular, 1.9–16.3 × 3–10.5 mm, 0.84–1.17(–2.53) times as long as wide, base rounded or convex, margins flat, entire, ciliate, apex retuse, rounded, convex, or acute, abaxial surface glabrous, adaxial highly glossy, glabrous; proximal blade margins entire; juvenile blade glabrous or puberulent. |
stipules foliaceous, rudimentary, or absent on early ones, foliaceous on late ones, apex acute; petiole convex to flat, or shallowly grooved adaxially, 4–9 mm, pilose adaxially; largest medial blade (sometimes amphistomatous), lorate, narrowly oblong, narrowly elliptic, or oblanceolate, 35–95 × 7.5–20 mm, 3.4–7.1 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex acute, acuminate, or convex, abaxial surface glaucous (sometimes obscured by hairs), sparsely to moderately densely long- to short-silky or pubescent, hairs (white, sometimes also ferruginous), straight or wavy, adaxial dull or slightly glossy, sparsely or moderately densely pubescent or long- to short-silky, (hairs white, sometimes also ferruginous); proximal blade margins entire or serrulate; juvenile blade densely long-silky abaxially, hairs white, sometimes also ferruginous. |
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Staminate flowers | abaxial nectary 0.5–1 mm, adaxial nectary narrowly oblong or oblong, 0.8–1.4 mm, nectaries distinct; filaments distinct or connate less than 1/2 their lengths, glabrous; anthers ellipsoid or globose, 0.4–0.6 mm. |
adaxial nectary oblong or ovate, 0.6–1.1 mm; filaments distinct or connate less than 1/2 their lengths, hairy basally; anthers purple turning yellow, 0.6–1 mm. |
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Pistillate flowers | abaxial nectary present or absent, adaxial nectary usually narrowly oblong or oblong, sometimes flask-shaped, 0.8–2 mm, longer than stipe; stipe 0.4–0.8 mm; ovary pyriform, glabrous or puberulent, (hairs in patches, especially on beak), beak slightly bulged below styles; ovules 7–17 per ovary; styles connate or slightly distinct distally, 0.5–1 mm; stigmas flat, abaxially non-papillate with pointed tip, or slenderly or broadly cylindrical, 0.28–0.6 mm. |
adaxial nectary oblong or flask-shaped, 0.7–1.3 mm, shorter than or equal to stipe; stipe 0.8–2 mm; ovary obclavate or pyriform, short-silky-villous, beak gradually tapering to or slightly bulged below styles; ovules 15–16 per ovary; styles 0.6–1 mm; stigmas flat, abaxially non-papillate with rounded or pointed tip, or broadly cylindrical, 0.24–0.32–0.44 mm. |
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Capsules | 3.8–8.3 mm. |
5–10 mm. |
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Catkins | from subterminal buds; staminate subglobose, stout, or indeterminate, 3.3–18.5 × 2.5–12 mm, flowering branchlet 0.5–9 mm; pistillate moderately densely to loosely flowered (2–15 flowers), stout, subglobose, globose, or indeterminate, 4.5–35 × 2–17 mm, flowering branchlet 0.5–22 mm; floral bract brown, 1.6–2.8 mm, apex rounded or retuse, entire, abaxially sparsely hairy or ciliate, hairs usually wavy, crinkled or curly, rarely straight. |
flowering as leaves emerge; staminate stout, 15.5–34 × 7–14 mm, flowering branchlet 1–8 mm; pistillate moderately densely flowered, stout, 20–55(–65 in fruit) × 11–13 mm, flowering branchlet 2–15 mm; floral bract dark brown or bicolor, 1.2–2.5 mm, apex acute or rounded, abaxially hairy, hairs straight or wavy. |
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Salix rotundifolia |
Salix orestera |
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Phenology | Flowering late May-late Aug. | |||||
Habitat | Subalpine meadows, slopes, lakes, streams, granite substrates | |||||
Elevation | 2100-4000 m (6900-13100 ft) | |||||
Distribution |
AK; MT; WY; NT; YT; e Asia
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CA; NV; OR
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Discussion | Varieties 2 (2 in the flora). Salix rotundifolia is closely related to S. polaris, from which it can be separated by its glabrous ovaries and fewer-flowered catkins. They also differ somewhat in leaf venation: S. rotundifolia typically having three main veins arising from the leaf base, often only one or two pair of secondary veins, and no or indistinct tertiary veins; S. polaris typically having pinnate venation, multiple secondary veins, and distinct tertiary veins. Salix rotundifolia consists of two varieties, the diploid var. dodgeana and the hexaploid var. rotundifolia. In general, var. dodgeana is a high alpine species in the southern cordillera of Wyoming and Montana, the St. Elias Mountains in Alaska and Yukon, the Mackenzie Mountains, Northwest Territories, and the Richardson Mountains, Yukon Territory. A diploid specimen of S. rotundifolia in the Cherski Mountains, Yakutia, Russia (B. A. Jurtzev and P. G. Zhukova 1982), which fits var. dodgeana in its 2–3-flowered catkins, relatively small leaves (3.5 × 3.9 mm), and small stomata (490 µm2), may represent an ancestral population. Variety rotundifolia usually occurs at lower elevations in Alaska and in easternmost Chukotka and Wrangel Island, Russia, but elevation separation is not distinct. There is a general correlation between stomatal size and ploidal level (W. Buechler, pers. comm.), but relatively large stomata in some diploid specimens of S. rotundifolia indicates a need for further cytological study. For the present, it is best to recognize the two cytotypes as varieties. Hybrids: Salix rotundifolia forms natural hybrids with S. arctica, S. phlebophylla, and S. polaris. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Salix orestera is so similar to S. glauca var. villosa that it is sometimes included in S. glauca. It is phenetically most similar to S. eastwoodiae and S. lemmonii (G. W. Argus 1997), and the possibility that it is a hybrid or an alloploid involving those species needs study. Salix orestera can be distinguished from S. glauca var. villosa in having largest medial blades usually narrower, 3.4–7.1 times as long as wide, abaxial surfaces distinctly silky with appressed hairs pointing toward apex, hairs sometimes ferruginous, secondary veins raised abaxially and adaxially, branches often strongly glaucous, and staminate flowers without abaxial nectaries; S. glauca var. villosa has largest medial blades usually broader, 2.2–3.9 times as long as wide, abaxial surfaces usually glabrescent or, if hairy, hairs unkempt, slightly spreading, hairs always white, secondary veins raised abaxially, flat or impressed adaxially, branches not glaucous, staminate flowers often with abaxial nectaries. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 69. | FNA vol. 7, p. 108. | ||||
Parent taxa | Salicaceae > Salix > subg. Chamaetia > sect. Herbella | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | S. commutata var. rubicunda, S. glauca subsp. orestera, S. glauca var. orestera | |||||
Name authority | Trautvetter: Nouv. Mém. Soc. Imp. Naturalistes Moscou 2: 304, plate 11. (1832) | C. K. Schneider: J. Arnold Arbor. 1: 164. (1920) | ||||
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