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net-leaf willow, net-vein willow, netted willow

Tyrrell's willow

Habit Plants 0.03–0.15 m, (dwarf, forming clones by layering). Shrubs, 0.6–3.5 m, (sometimes forming clones by layering).
Stems

trailing;

branches and branchlets yellow-brown or red-brown, glabrous.

branches and branchlets red-brown, not to strongly glaucous, glabrous, (buds caprea-type or intermediate).

Leaves

stipules absent or rudimentary;

petiole 3–46 mm, (sometimes glandular distally);

largest medial blade amphistomatous or hemiamphistomatous, (2 pairs of secondary veins arising at or close to base, arcing toward apex), oblong, broadly oblong, broadly elliptic, subcircular, or circular, (8–)12–66 × 8–50 mm, 1–1.5 times as long as wide, base convex, rounded, subcordate, or cordate, margins slightly revolute, entire or crenulate (glandular-dotted), apex rounded, convex, or retuse, abaxial surface sparsely long-silky to glabrescent, adaxial (venation deeply impressed), slightly or highly glossy, glabrous or pilose;

proximal blade margins entire;

juvenile blade glabrous.

stipules rudimentary or foliaceous (often brownish) on early ones, foliaceous or rudimentary on late ones, (ca. 4 mm), apex acute;

petiole convex to flat, or shallowly grooved adaxially, 1–3.4–16 mm, glabrous, pilose, or short-silky adaxially;

largest medial blade (amphistomatous), narrowly elliptic, elliptic, oblanceolate, or obovate, 15–29–65 × 3.5–8.8–18 mm, 2.3–3.3–4.4 times as long as wide, base cuneate or convex, margins strongly to slightly revolute, entire, or very shallowly serrulate or shallowly serrulate-crenulate, apex acute, acuminate, or convex, abaxial surface glaucous, sparsely long-silky to glabrescent, hairs (ferruginous), straight, adaxial highly glossy, glabrous or sparsely short-silky (hairs white, sometimes also ferruginous);

proximal blade margins entire;

juvenile blade yellowish green or sometimes reddish, sparsely long-silky abaxially, hairs white and ferruginous.

Staminate flowers

abaxial nectary 0.5–0.9 mm, adaxial nectary oblong or ovate, 0.5–1 mm, nectaries connate and cup-shaped;

filaments distinct, hairy on proximal 1/2 or throughout;

anthers ellipsoid or globose, 0.3–0.4 mm.

adaxial nectary oblong, 0.8–1.1 mm;

filaments distinct, glabrous or hairy basally;

anthers purple turning yellow, ellipsoid, 0.4–0.7 mm.

Pistillate flowers

abaxial nectary (0–)0.3–0.5 mm, adaxial nectary narrowly oblong, 0.5–1 mm, equal to or longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.8 mm;

ovary pyriform or ovoid, short-silky, hairs flattened, beak abruptly tapering to styles;

ovules 8–18 per ovary;

styles connate to distinct 1/2 their lengths, 0.2–0.3 mm;

stigmas flat, abaxially non-papillate with rounded tip, broadly cylindrical, or 2 plump lobes, 0.2–0.26–0.36 mm.

adaxial nectary oblong or flask-shaped, 0.6–1.1 mm, equal to or shorter than stipe;

stipe 0.2–1 mm (–1.4 mm in cultivation);

ovary pyriform, long-silky, beak gradually tapering to styles;

ovules 12–16 per ovary;

styles 0.6–1.2 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.44–0.55–0.75 mm.

Capsules

4.5–5 mm.

3.6–5 mm.

Catkins

staminate 11–54 × 4–9 mm, flowering branchlet 2–28 mm; pistillate densely flowered (more than 6 flowers), slender or stout, 11–79 × 3–8 mm, flowering branchlet 2–37 mm;

floral bract tawny, 0.8–1.8 mm, apex rounded to retuse, entire, abaxially glabrous.

flowering before leaves emerge; staminate stout, 14–35 × 12–16 mm, flowering branchlet 0 mm; pistillate densely flowered, stout, 17–51 × 10–13–22 mm, flowering branchlet 0–4 mm;

floral bract brown, black, or bicolor, 1–3.7 mm, apex acute to acuminate or rounded, abaxially hairy, hairs long, straight.

2n

= 38.

Salix reticulata

Salix tyrrellii

Phenology Flowering early Jun-mid Aug. Flowering mid Jun-mid Jul.
Habitat Arctic-alpine, polygonal tundra, dry tussock tundra, partially stabilized sand dunes, sedge meadows, Dryas tundra on alpine cliffs and ledges, snowbeds, stabilized talus slopes, white spruce woods, treed bogs Active sand dunes, shrubby tundra
Elevation 0-3500 m (0-11500 ft) 200-600 m (700-2000 ft)
Distribution
from FNA
AK; CO; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Europe; Asia (Chukotka, Russian Far East, arctic, e Siberia, Spitzbergen)
[WildflowerSearch map]
[BONAP county map]
from FNA
AB; NT; NU; SK
[BONAP county map]
Discussion

The reported occurrence of Salix reticulata in Colorado (R. D. Dorn 1997) needs further study.

Salix reticulata occurs in Europe in northern Scotland, northern Scandinavia, the Alps and other European mountains, and arctic Eurasia. The species is circumpolar except for Greenland and Iceland.

A population of Salix reticulata on the Queen Charlotte Islands, with consistently glabrous ovaries, was named subsp. glabellicarpa. Some southeastern Alaska populations have plants with glabrous, partially hairy, and completely short-silky ovaries growing together. The possibility that subsp. glabellicarpa may be a hybrid or a simple mutation needs study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix tyrrellii, first described from Lake Athabasca sand dunes in northwestern Saskatchewan and adjacent Alberta, is characterized by slender, amphistomatous leaves and relatively long, slender branchlets (G. W. Argus and J. W. Steele 1979). It probably evolved, in the past 10,000 years, from the widespread boreal S. planifolia. Originally, it was thought to be endemic to Lake Athabasca but recent collections from Nunavut and the Northwest Territories suggest that it may have a much wider range. Plants from Nunavut and the Northwest Territories do not have the long, slender branchlets found in sand dune populations, and identification is primarily based on the presence of amphistomatous leaves. This character, however, may be unreliable. For example, S. planifolia from 2610 m in the Glass Mountains, California, have amphistomatous leaves, suggesting that this character may be under environmental influence. The appropriate rank for this taxon remains uncertain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 65. FNA vol. 7, p. 140.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Vetrix > sect. Phylicifoliae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. reticulata var. gigantifolia, S. reticulata subsp. glabellicarpa, S. reticulata var. semicalva S. planifolia subsp. tyrrellii
Name authority Linnaeus: Sp. Pl. 2: 1018. (1753) Raup: J. Arnold Arbor. 17: 231, plate 192. (1936)
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