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net-leaf willow, net-vein willow, netted willow

Tracy's willow

Habit Plants 0.03–0.15 m, (dwarf, forming clones by layering). Shrubs, 1–6 m. Stems: branches gray-brown to red-brown, strongly to weakly glaucous, glabrous; branchlets yellow-brown to red-brown, (weakly glaucous), glabrous or sparsely to moderately densely velvety or tomentose.
Stems

trailing;

branches and branchlets yellow-brown or red-brown, glabrous.

Leaves

stipules absent or rudimentary;

petiole 3–46 mm, (sometimes glandular distally);

largest medial blade amphistomatous or hemiamphistomatous, (2 pairs of secondary veins arising at or close to base, arcing toward apex), oblong, broadly oblong, broadly elliptic, subcircular, or circular, (8–)12–66 × 8–50 mm, 1–1.5 times as long as wide, base convex, rounded, subcordate, or cordate, margins slightly revolute, entire or crenulate (glandular-dotted), apex rounded, convex, or retuse, abaxial surface sparsely long-silky to glabrescent, adaxial (venation deeply impressed), slightly or highly glossy, glabrous or pilose;

proximal blade margins entire;

juvenile blade glabrous.

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones, (2.5–9 mm), apex rounded or convex;

petiole convex to flat, or shallowly grooved adaxially, 5–11 mm, tomentose or velvety adaxially;

largest medial blade lorate, oblanceolate, or elliptic, 55–96 × 15–34 mm, 2–3.7 times as long as wide, base convex, margins slightly revolute, entire, sinuate or serrulate, apex acuminate to acute, abaxial surface glaucous, sparsely pubescent to moderately densely tomentose or glabrous, hairs (white, sometimes also ferruginous), straight or curved, adaxial slightly glossy, sparsely tomentose to glabrescent;

proximal blade margins entire or serrulate;

juvenile blade reddish, very densely tomentose or puberulent to glabrescent abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

abaxial nectary 0.5–0.9 mm, adaxial nectary oblong or ovate, 0.5–1 mm, nectaries connate and cup-shaped;

filaments distinct, hairy on proximal 1/2 or throughout;

anthers ellipsoid or globose, 0.3–0.4 mm.

adaxial nectary narrowly oblong, oblong, or ovate, 0.3–0.8 mm;

filaments connate less to more than 1/2 their lengths;

anthers purple turning yellow, ellipsoid, shortly cylindrical, or globose, 0.36–0.44 mm.

Pistillate flowers

abaxial nectary (0–)0.3–0.5 mm, adaxial nectary narrowly oblong, 0.5–1 mm, equal to or longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.8 mm;

ovary pyriform or ovoid, short-silky, hairs flattened, beak abruptly tapering to styles;

ovules 8–18 per ovary;

styles connate to distinct 1/2 their lengths, 0.2–0.3 mm;

stigmas flat, abaxially non-papillate with rounded tip, broadly cylindrical, or 2 plump lobes, 0.2–0.26–0.36 mm.

adaxial nectary oblong, or narrowly oblong to flask-shaped, 0.2–0.6 mm;

stipe 1–1.7(–2.4) mm;

ovary pyriform, beak sometimes slightly bulged below styles;

ovules 12 per ovary;

styles 0.1–0.6 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.1–0.3 mm.

Capsules

4.5–5 mm.

2.4–3.6 mm.

Catkins

staminate 11–54 × 4–9 mm, flowering branchlet 2–28 mm; pistillate densely flowered (more than 6 flowers), slender or stout, 11–79 × 3–8 mm, flowering branchlet 2–37 mm;

floral bract tawny, 0.8–1.8 mm, apex rounded to retuse, entire, abaxially glabrous.

flowering as or just before leaves emerge; staminate stout, 17–30 × 7–9 mm, flowering branchlet 1.5–5 mm; pistillate very densely flowered, stout to slender, 17–42 × 6–11 mm, flowering branchlet 1.5–3 mm;

floral bract brown or bicolor, 0.8–1.6 mm, apex rounded to truncate, abaxially hairy throughout or proximally, hairs wavy or straight.

2n

= 38.

Salix reticulata

Salix tracyi

Phenology Flowering early Jun-mid Aug. Flowering early Apr-early May.
Habitat Arctic-alpine, polygonal tundra, dry tussock tundra, partially stabilized sand dunes, sedge meadows, Dryas tundra on alpine cliffs and ledges, snowbeds, stabilized talus slopes, white spruce woods, treed bogs Shores and floodplains, sandy, gravelly, or rocky substrates, often serpentine
Elevation 0-3500 m (0-11500 ft) 90-500 m (300-1600 ft)
Distribution
from FNA
AK; CO; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Europe; Asia (Chukotka, Russian Far East, arctic, e Siberia, Spitzbergen)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

The reported occurrence of Salix reticulata in Colorado (R. D. Dorn 1997) needs further study.

Salix reticulata occurs in Europe in northern Scotland, northern Scandinavia, the Alps and other European mountains, and arctic Eurasia. The species is circumpolar except for Greenland and Iceland.

A population of Salix reticulata on the Queen Charlotte Islands, with consistently glabrous ovaries, was named subsp. glabellicarpa. Some southeastern Alaska populations have plants with glabrous, partially hairy, and completely short-silky ovaries growing together. The possibility that subsp. glabellicarpa may be a hybrid or a simple mutation needs study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ball described Salix tracyi as part of a group of “more or less localized endemic willows of northern California and southern Oregon.” A. K. Skvortsov (1971), followed by G. W. Argus (1993), thought that this species was a stunted form, perhaps shade-growth, of S. lasiolepis. R. D. Dorn (2000) studied the taxon in the field and considered it to be a distinct species. He noted that it could “usually” be separated from S. lasiolepis by its shorter styles, longer stipes, less hairy floral bracts, shorter catkins, thinner, less hairy leaves, and reddish juvenile leaves. He also noted differences in flowering times but they overlap completely. The differences are subtle and, although not diagnostic, should be studied further. Salix tracyi is treated here as a species because of its relatively large, localized range and interesting biological and morphological characteristics. Local botanists making collections from tagged individuals could provide useful information on habitat, elevation, flowering time, and morphology. Experimental study of this and the other serpentine endemic Salix in the region (see also 110. S. breweri and 111. S. delnortensis) could provide insights into their evolution and their relationship with S. lasiolepis.

A numerical taxonomic study by T. J. Crovello (1968) found that Salix tracyi clustered with members of sect. Cordatae, but both G. W. Argus (1997) and R. D. Dorn (2000) placed it in sect. Mexicanae with S. lasiolepis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 65. FNA vol. 7, p. 156.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Vetrix > sect. Mexicanae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. reticulata var. gigantifolia, S. reticulata subsp. glabellicarpa, S. reticulata var. semicalva
Name authority Linnaeus: Sp. Pl. 2: 1018. (1753) C. R. Ball: Univ. Calif. Publ. Bot. 17: 403, plates 69, 70. (1934)
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