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net-leaf willow, net-vein willow, netted willow

Richardson's willow

Habit Plants 0.03–0.15 m, (dwarf, forming clones by layering). Plants 0.3–6.5 m. Stems: branches red-brown, yellow-brown, or violet, not to strongly glaucous (slightly or highly glossy), villous to glabrescent with persistent patches of hairs; branchlets red-brown or yellow-brown, pilose to densely villous, (buds caprea-type, scale inner membrane free, separating from outer layer).
Stems

trailing;

branches and branchlets yellow-brown or red-brown, glabrous.

Leaves

stipules absent or rudimentary;

petiole 3–46 mm, (sometimes glandular distally);

largest medial blade amphistomatous or hemiamphistomatous, (2 pairs of secondary veins arising at or close to base, arcing toward apex), oblong, broadly oblong, broadly elliptic, subcircular, or circular, (8–)12–66 × 8–50 mm, 1–1.5 times as long as wide, base convex, rounded, subcordate, or cordate, margins slightly revolute, entire or crenulate (glandular-dotted), apex rounded, convex, or retuse, abaxial surface sparsely long-silky to glabrescent, adaxial (venation deeply impressed), slightly or highly glossy, glabrous or pilose;

proximal blade margins entire;

juvenile blade glabrous.

stipules (sometimes marcescent), foliaceous, (3–12–35 mm, 1.7–3.6–6.6 times as long as wide, lanceolate or narrowly elliptic, sometimes ovate or oval, base broad, abruptly tapering), apex acuminate or acute;

petiole convex to flat or shallowly grooved adaxially, 2–7.3–27 mm, villous to glabrescent adaxially;

largest medial blade broadly to narrowly elliptic, or obovate, 23–45–100 × 10–22–55 mm, 1.2–2.2–4.2 times as long as wide, base cuneate, convex, or concave, margins slightly revolute or flat, entire or serrulate, apex acute, acuminate, or convex, abaxial surface glaucous, glabrous or pilose, hairs (white, sometimes also ferruginous), curved, adaxial slightly glossy, moderately densely villous or pilose to glabrescent;

proximal blade margins usually entire, sometimes serrulate;

juvenile blade yellowish green, abaxially densely villous or pilose, sometimes glabrous, hairs white, sometimes also ferruginous.

Staminate flowers

abaxial nectary 0.5–0.9 mm, adaxial nectary oblong or ovate, 0.5–1 mm, nectaries connate and cup-shaped;

filaments distinct, hairy on proximal 1/2 or throughout;

anthers ellipsoid or globose, 0.3–0.4 mm.

adaxial nectary narrowly oblong to oblong, 0.7–1.8 mm;

filaments distinct to connate ca. 1/2 their lengths, glabrous;

anthers purple turning yellow, ellipsoid to shortly cylindrical, 0.5–0.8 mm.

Pistillate flowers

abaxial nectary (0–)0.3–0.5 mm, adaxial nectary narrowly oblong, 0.5–1 mm, equal to or longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.8 mm;

ovary pyriform or ovoid, short-silky, hairs flattened, beak abruptly tapering to styles;

ovules 8–18 per ovary;

styles connate to distinct 1/2 their lengths, 0.2–0.3 mm;

stigmas flat, abaxially non-papillate with rounded tip, broadly cylindrical, or 2 plump lobes, 0.2–0.26–0.36 mm.

adaxial nectary narrowly oblong, oblong, or flask-shaped, 0.6–1.4 mm (2+-lobed), longer than stipe;

stipe 0.4–1 mm;

ovary pyriform, beak gradually tapering to styles;

ovules 22–37 per ovary;

styles 1–2.8 mm;

stigmas slenderly cylindrical, 0.32–0.63–0.86 mm.

Capsules

4.5–5 mm.

4.5–6.8 mm.

Catkins

staminate 11–54 × 4–9 mm, flowering branchlet 2–28 mm; pistillate densely flowered (more than 6 flowers), slender or stout, 11–79 × 3–8 mm, flowering branchlet 2–37 mm;

floral bract tawny, 0.8–1.8 mm, apex rounded to retuse, entire, abaxially glabrous.

flowering before leaves emerge; staminate stout or subglobose, 22–53 × 16–24 mm, flowering branchlet 0 mm; pistillate densely flowered, slender or stout, 25–69(–85 in fruit) × 13–20 mm, flowering branchlet 0–2 mm;

floral bract brown or black, 2.4–3.2 mm, apex acute, rounded, or retuse, abaxially moderately densely hairy, hairs straight.

2n

= 38.

= 38.

Salix reticulata

Salix richardsonii

Phenology Flowering early Jun-mid Aug. Flowering late May-early Jul.
Habitat Arctic-alpine, polygonal tundra, dry tussock tundra, partially stabilized sand dunes, sedge meadows, Dryas tundra on alpine cliffs and ledges, snowbeds, stabilized talus slopes, white spruce woods, treed bogs Arctic, subarctic, subalpine and boreal, stream terraces and floodplains, open forests, wet sedge meadows, fens, drumlin fields, gravel ridges, bogs, cliff ledges, snowbeds, silt, sand and gravel, calcareous or igneous substrates
Elevation 0-3500 m (0-11500 ft) 10-1800 m (0-5900 ft)
Distribution
from FNA
AK; CO; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Europe; Asia (Chukotka, Russian Far East, arctic, e Siberia, Spitzbergen)
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC; MB; NT; NU; YT; e Asia (Chukotka, Kamchatka, Russia [Anadyr], e Siberia)
[BONAP county map]
Discussion

The reported occurrence of Salix reticulata in Colorado (R. D. Dorn 1997) needs further study.

Salix reticulata occurs in Europe in northern Scotland, northern Scandinavia, the Alps and other European mountains, and arctic Eurasia. The species is circumpolar except for Greenland and Iceland.

A population of Salix reticulata on the Queen Charlotte Islands, with consistently glabrous ovaries, was named subsp. glabellicarpa. Some southeastern Alaska populations have plants with glabrous, partially hairy, and completely short-silky ovaries growing together. The possibility that subsp. glabellicarpa may be a hybrid or a simple mutation needs study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybrids:

Salix richardsonii forms natural hybrids with S. barclayi and S. calcicola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 65. FNA vol. 7, p. 144.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Vetrix > sect. Lanatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. reticulata var. gigantifolia, S. reticulata subsp. glabellicarpa, S. reticulata var. semicalva S. lanata subsp. richardsonii, S. richardsonii var. mckeandii
Name authority Linnaeus: Sp. Pl. 2: 1018. (1753) Hooker: Fl. Bor.-Amer. 2: 147, plate 182. (1838)
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