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net-leaf willow, net-vein willow, netted willow

meadow willow, skeleton-leaf willow, slender willow

Habit Plants 0.03–0.15 m, (dwarf, forming clones by layering). Plants 1–6 m. Stems: branches red-brown or violet, not or weakly glaucous, (dull or slightly glossy), puberulent; branchlets yellow-green to red-brown, sparsely pubescent or moderately densely velvety, (buds alba-type or intermediate).
Stems

trailing;

branches and branchlets yellow-brown or red-brown, glabrous.

Leaves

stipules absent or rudimentary;

petiole 3–46 mm, (sometimes glandular distally);

largest medial blade amphistomatous or hemiamphistomatous, (2 pairs of secondary veins arising at or close to base, arcing toward apex), oblong, broadly oblong, broadly elliptic, subcircular, or circular, (8–)12–66 × 8–50 mm, 1–1.5 times as long as wide, base convex, rounded, subcordate, or cordate, margins slightly revolute, entire or crenulate (glandular-dotted), apex rounded, convex, or retuse, abaxial surface sparsely long-silky to glabrescent, adaxial (venation deeply impressed), slightly or highly glossy, glabrous or pilose;

proximal blade margins entire;

juvenile blade glabrous.

stipules rudimentary or absent;

petiole shallowly grooved adaxially, 3–11 mm, pubescent, or velvety to glabrescent adaxially;

largest medial blade lorate or very narrowly elliptic, 38–110 × 6–19 mm, 5–9 times as long as wide, base cuneate or convex, margins flat to slightly revolute, entire, serrate, serrulate, or spinulose-serrate, apex acute to acuminate, abaxial surface glaucous, densely long-silky to glabrescent, hairs (white, sometimes also ferruginous), adaxial dull or slightly glossy, glabrous or sparsely pubescent, (hairs white, sometimes also ferruginous);

proximal blade margins sometimes serrulate;

juvenile blade moderately densely long-silky abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

abaxial nectary 0.5–0.9 mm, adaxial nectary oblong or ovate, 0.5–1 mm, nectaries connate and cup-shaped;

filaments distinct, hairy on proximal 1/2 or throughout;

anthers ellipsoid or globose, 0.3–0.4 mm.

adaxial nectary square, ovate, or oblong, 0.3–0.7 mm;

filaments distinct, hairy basally;

anthers purple turning yellow, ellipsoid or globose, 0.4–0.6 mm.

Pistillate flowers

abaxial nectary (0–)0.3–0.5 mm, adaxial nectary narrowly oblong, 0.5–1 mm, equal to or longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.8 mm;

ovary pyriform or ovoid, short-silky, hairs flattened, beak abruptly tapering to styles;

ovules 8–18 per ovary;

styles connate to distinct 1/2 their lengths, 0.2–0.3 mm;

stigmas flat, abaxially non-papillate with rounded tip, broadly cylindrical, or 2 plump lobes, 0.2–0.26–0.36 mm.

adaxial nectary oblong to ovate, 0.3–0.9 mm;

stipe 1.5–4 mm;

ovary pyriform, beak abruptly tapering to styles;

ovules 6–12 per ovary;

styles 0–0.5 mm;

stigmas slenderly to broadly cylindrical, 0.26–0.4–0.8 mm.

Capsules

4.5–5 mm.

5–9 mm.

Catkins

staminate 11–54 × 4–9 mm, flowering branchlet 2–28 mm; pistillate densely flowered (more than 6 flowers), slender or stout, 11–79 × 3–8 mm, flowering branchlet 2–37 mm;

floral bract tawny, 0.8–1.8 mm, apex rounded to retuse, entire, abaxially glabrous.

flowering as leaves emerge; staminate stout to globose, 12–29 × 6–17 mm, flowering branchlet 0.8–3 mm; pistillate loosely flowered, stout to globose, 12–39 × 6–18 mm, flowering branchlet 1–11 mm;

floral bract brown, tawny, light rose, or bicolor, 1–2 mm, apex rounded, abaxially sparsely hairy, hairs straight.

2n

= 38.

= 38.

Salix reticulata

Salix petiolaris

Phenology Flowering early Jun-mid Aug. Flowering mid Apr-mid Jun.
Habitat Arctic-alpine, polygonal tundra, dry tussock tundra, partially stabilized sand dunes, sedge meadows, Dryas tundra on alpine cliffs and ledges, snowbeds, stabilized talus slopes, white spruce woods, treed bogs Sedge meadows, openings in moist, low, rich deciduous woods, sandy or peaty wet prairies, lakeshores
Elevation 0-3500 m (0-11500 ft) 10-2700 m (0-8900 ft)
Distribution
from FNA
AK; CO; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Europe; Asia (Chukotka, Russian Far East, arctic, e Siberia, Spitzbergen)
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; CT; IA; IL; IN; MA; ME; MI; MN; ND; NE; NH; NJ; NY; OH; PA; SD; VT; WI; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

The reported occurrence of Salix reticulata in Colorado (R. D. Dorn 1997) needs further study.

Salix reticulata occurs in Europe in northern Scotland, northern Scandinavia, the Alps and other European mountains, and arctic Eurasia. The species is circumpolar except for Greenland and Iceland.

A population of Salix reticulata on the Queen Charlotte Islands, with consistently glabrous ovaries, was named subsp. glabellicarpa. Some southeastern Alaska populations have plants with glabrous, partially hairy, and completely short-silky ovaries growing together. The possibility that subsp. glabellicarpa may be a hybrid or a simple mutation needs study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

See 107. Salix sericea for a comparative description.

Because reproductive barriers between Salix petiolaris and S. eriocephala are weak, A. Mosseler (1990) suggested that their morphological variability may be due to interspecific gene flow.

Hybrids:

Salix petiolaris forms natural hybrids with S. bebbiana, S. candida, S. eriocephala, S. famelica, S. pellita, and S. sericea. Hybrids with S. alba have been reported (M. L. Fernald 1950) but no convincing specimens have been seen. Controlled pollinations with S. discolor produced no seed (A. Mosseler 1990).

Reports of Salix petiolaris × S. sericea from Massachusetts and Pennsylvania (C. K. Schneider 1921) probably refer to the densely sericeous variant of S. petiolaris. It is sometimes named S. ×subsericea (Andersson) C. K. Schneider but does not seem to be a hybrid (G. W. Argus 1965, 1986; E. G. Voss 1972–1996, vol. 2).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 65. FNA vol. 7, p. 152.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Vetrix > sect. Geyerianae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. reticulata var. gigantifolia, S. reticulata subsp. glabellicarpa, S. reticulata var. semicalva S. gracilis, S. gracilis var. textoris, S. petiolaris var. gracilis, S. ×subsericea
Name authority Linnaeus: Sp. Pl. 2: 1018. (1753) Smith: Trans. Linn. Soc. 6: 122. (1802)
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