Salix reticulata |
Salix humilis |
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net-leaf willow, net-vein willow, netted willow |
gray or small pussy or upland willow, prairie willow |
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Habit | Plants 0.03–0.15 m, (dwarf, forming clones by layering). | Shrubs, 0.3–3 m, (forming clones by layering). | ||||
Stems | trailing; branches and branchlets yellow-brown or red-brown, glabrous. |
branches dark red-brown, not or weakly glaucous, tomentose to glabrescent; branchlets red-brown, yellow-brown, or greenish brown, moderately to very densely villous, tomentose, or velvety-tomentose to glabrescent. |
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Leaves | stipules absent or rudimentary; petiole 3–46 mm, (sometimes glandular distally); largest medial blade amphistomatous or hemiamphistomatous, (2 pairs of secondary veins arising at or close to base, arcing toward apex), oblong, broadly oblong, broadly elliptic, subcircular, or circular, (8–)12–66 × 8–50 mm, 1–1.5 times as long as wide, base convex, rounded, subcordate, or cordate, margins slightly revolute, entire or crenulate (glandular-dotted), apex rounded, convex, or retuse, abaxial surface sparsely long-silky to glabrescent, adaxial (venation deeply impressed), slightly or highly glossy, glabrous or pilose; proximal blade margins entire; juvenile blade glabrous. |
stipules absent or rudimentary on early ones, absent, rudimentary or foliaceous on late ones, apex acute; petiole convex to flat, or shallowly grooved adaxially, 0.5–7(–12) mm, velvety, pilose, or villous adaxially; largest medial blade (sometimes hemiamphistomatous), narrowly oblong, narrowly elliptic, elliptic, oblanceolate, obovate, or broadly obovate, (13–)20–90(–135) × 3–23(–35) mm, 2.3–9 times as long as wide, base cuneate or convex, margins revolute or flat, entire, crenate, or sinuate, (glands submarginal), apex acuminate or convex, abaxial surface glaucous, sparsely to densely tomentose or woolly, hairs erect or spreading, wavy, adaxial slightly or highly glossy, glabrous, pubescent, tomentose, or pilose; proximal blade margins entire or serrulate; juvenile blade green, densely tomentose to glabrescent abaxially, hairs white, sometimes also ferruginous. |
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Staminate flowers | abaxial nectary 0.5–0.9 mm, adaxial nectary oblong or ovate, 0.5–1 mm, nectaries connate and cup-shaped; filaments distinct, hairy on proximal 1/2 or throughout; anthers ellipsoid or globose, 0.3–0.4 mm. |
adaxial nectary oblong or square, 0.2–0.7 mm; filaments distinct, glabrous or hairy basally; anthers purple turning yellow, ellipsoid or cylindrical, 0.4–0.6 mm. |
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Pistillate flowers | abaxial nectary (0–)0.3–0.5 mm, adaxial nectary narrowly oblong, 0.5–1 mm, equal to or longer than stipe, nectaries distinct or connate and cup-shaped; stipe 0–0.8 mm; ovary pyriform or ovoid, short-silky, hairs flattened, beak abruptly tapering to styles; ovules 8–18 per ovary; styles connate to distinct 1/2 their lengths, 0.2–0.3 mm; stigmas flat, abaxially non-papillate with rounded tip, broadly cylindrical, or 2 plump lobes, 0.2–0.26–0.36 mm. |
adaxial nectary square, 0.4–0.8 mm, shorter than stipe; stipe 1–2.5 mm; ovary obclavate or pyriform, moderately densely to sparsely short-silky-villous (hairs refractive), beak slightly bulged below styles, (valves recurving in fruit); ovules 6–12 per ovary; styles (sometimes slightly distinct distally), 0.2–0.4 mm; stigmas slenderly to broadly cylindrical, 0.2–0.56 mm. |
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Capsules | 4.5–5 mm. |
5–12 mm. |
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Catkins | staminate 11–54 × 4–9 mm, flowering branchlet 2–28 mm; pistillate densely flowered (more than 6 flowers), slender or stout, 11–79 × 3–8 mm, flowering branchlet 2–37 mm; floral bract tawny, 0.8–1.8 mm, apex rounded to retuse, entire, abaxially glabrous. |
flowering before leaves emerge; staminate 6.5–34 × 5–19 mm, flowering branchlet 0–1 mm; pistillate (and staminate) moderately to very densely flowered, stout, subglobose, or globose, 9–47(–55 in fruit) × 5.5–19 mm, flowering branchlet 0–4 mm; floral bract brown, black, or bicolor, 0.8–2 mm, apex rounded or acute, abaxially moderately densely hairy, hairs (white), straight or wavy. |
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2n | = 38. |
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Salix reticulata |
Salix humilis |
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Phenology | Flowering early Jun-mid Aug. | |||||
Habitat | Arctic-alpine, polygonal tundra, dry tussock tundra, partially stabilized sand dunes, sedge meadows, Dryas tundra on alpine cliffs and ledges, snowbeds, stabilized talus slopes, white spruce woods, treed bogs | |||||
Elevation | 0-3500 m (0-11500 ft) | |||||
Distribution |
AK; CO; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Europe; Asia (Chukotka, Russian Far East, arctic, e Siberia, Spitzbergen)
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; WI; WV; MB; NB; NS; ON; PE; QC
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Discussion | The reported occurrence of Salix reticulata in Colorado (R. D. Dorn 1997) needs further study. Salix reticulata occurs in Europe in northern Scotland, northern Scandinavia, the Alps and other European mountains, and arctic Eurasia. The species is circumpolar except for Greenland and Iceland. A population of Salix reticulata on the Queen Charlotte Islands, with consistently glabrous ovaries, was named subsp. glabellicarpa. Some southeastern Alaska populations have plants with glabrous, partially hairy, and completely short-silky ovaries growing together. The possibility that subsp. glabellicarpa may be a hybrid or a simple mutation needs study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Varieties humilis and tristis are almost sympatric. In Maine they occur in the same populations without evident intermediate forms, while in other places, e.g., Florida, Georgia, and Nebraska, they apparently intergrade. Usually, they are distinct in habit, leaf and catkin size, and, most conspicuously, the presence of foliaceous stipules in var. humilis and their absence in var. tristis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 65. | FNA vol. 7, p. 129. | ||||
Parent taxa | Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia | Salicaceae > Salix > subg. Vetrix > sect. Cinerella | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | S. reticulata var. gigantifolia, S. reticulata subsp. glabellicarpa, S. reticulata var. semicalva | |||||
Name authority | Linnaeus: Sp. Pl. 2: 1018. (1753) | Marshall: Arbust. Amer., 140. (1785) | ||||
Web links |