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net-leaf willow, net-vein willow, netted willow

hoary willow, sage willow, sage-leaf willow

Habit Plants 0.03–0.15 m, (dwarf, forming clones by layering). Plants often forming clones by layering.
Stems

trailing;

branches and branchlets yellow-brown or red-brown, glabrous.

branches dark gray-brown to yellow-brown, not glaucous, woolly in patches or floccose to glabrescent;

branchlets yellow-brown to red-brown or gray-brown, densely (white) woolly or tomentose, sometimes floccose.

Leaves

stipules absent or rudimentary;

petiole 3–46 mm, (sometimes glandular distally);

largest medial blade amphistomatous or hemiamphistomatous, (2 pairs of secondary veins arising at or close to base, arcing toward apex), oblong, broadly oblong, broadly elliptic, subcircular, or circular, (8–)12–66 × 8–50 mm, 1–1.5 times as long as wide, base convex, rounded, subcordate, or cordate, margins slightly revolute, entire or crenulate (glandular-dotted), apex rounded, convex, or retuse, abaxial surface sparsely long-silky to glabrescent, adaxial (venation deeply impressed), slightly or highly glossy, glabrous or pilose;

proximal blade margins entire;

juvenile blade glabrous.

stipules rudimentary or foliaceous on early ones, late ones 2–3.6 mm, apex acute;

petiole shallowly to deeply grooved adaxially, 3–10 mm, tomentose or densely (white) woolly adaxially (obscured by hairs);

largest medial blade lorate, narrowly elliptic or oblanceolate, 47–103 × 5–20 mm, base convex or cuneate, margins strongly to slightly revolute, entire, or sinuate, apex acute or convex, abaxial surface glaucous (generally obscured by hairs), very densely to sparsely tomentose-woolly (cobwebby in age), hairs dull white, crinkled, adaxial dull or slightly glossy, moderately densely to sparsely tomentose, floccose, hairs dull white;

proximal blade margins entire;

juvenile blade yellowish green, very densely tomentose abaxially, hairs white.

Staminate flowers

abaxial nectary 0.5–0.9 mm, adaxial nectary oblong or ovate, 0.5–1 mm, nectaries connate and cup-shaped;

filaments distinct, hairy on proximal 1/2 or throughout;

anthers ellipsoid or globose, 0.3–0.4 mm.

adaxial nectary narrowly oblong to oblong, 0.6–1 mm;

filaments distinct or connate less than 1/2 their lengths;

anthers purple turning yellow, ellipsoid, long-cylindrical, or globose, 0.4–0.6 mm.

Pistillate flowers

abaxial nectary (0–)0.3–0.5 mm, adaxial nectary narrowly oblong, 0.5–1 mm, equal to or longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.8 mm;

ovary pyriform or ovoid, short-silky, hairs flattened, beak abruptly tapering to styles;

ovules 8–18 per ovary;

styles connate to distinct 1/2 their lengths, 0.2–0.3 mm;

stigmas flat, abaxially non-papillate with rounded tip, broadly cylindrical, or 2 plump lobes, 0.2–0.26–0.36 mm.

adaxial nectary oblong, 0.4–1 mm;

ovary pyriform, beak sometimes slightly bulged below styles;

ovules 12–18 per ovary;

styles 0.3–1.9 mm.

Capsules

4.5–5 mm.

4–6 mm.

Catkins

staminate 11–54 × 4–9 mm, flowering branchlet 2–28 mm; pistillate densely flowered (more than 6 flowers), slender or stout, 11–79 × 3–8 mm, flowering branchlet 2–37 mm;

floral bract tawny, 0.8–1.8 mm, apex rounded to retuse, entire, abaxially glabrous.

flowering as leaves emerge; staminate stout or subglobose, 17–39 × 8–16 mm, flowering branchlet 0.5–7 mm; pistillate densely to moderately densely flowered, stout or slender, 20–66 × 9–18 mm, flowering branchlet 1–24 mm;

floral bract tawny or brown, 1.2–1.8 mm, apex rounded or acute, abaxially hairy, hairs straight to wavy.

2n

= 38.

= 38.

Salix reticulata

Salix candida

Phenology Flowering early Jun-mid Aug. Flowering mid Apr-early Jul.
Habitat Arctic-alpine, polygonal tundra, dry tussock tundra, partially stabilized sand dunes, sedge meadows, Dryas tundra on alpine cliffs and ledges, snowbeds, stabilized talus slopes, white spruce woods, treed bogs Floodplains, marl bogs, fens, and meadows, calcareous substrates
Elevation 0-3500 m (0-11500 ft) 10-2800 m (0-9200 ft)
Distribution
from FNA
AK; CO; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT; Europe; Asia (Chukotka, Russian Far East, arctic, e Siberia, Spitzbergen)
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; CT; IA; ID; IL; IN; MA; ME; MI; MN; MT; ND; NH; NJ; NY; OH; PA; SD; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; SPM
[WildflowerSearch map]
[BONAP county map]
Discussion

The reported occurrence of Salix reticulata in Colorado (R. D. Dorn 1997) needs further study.

Salix reticulata occurs in Europe in northern Scotland, northern Scandinavia, the Alps and other European mountains, and arctic Eurasia. The species is circumpolar except for Greenland and Iceland.

A population of Salix reticulata on the Queen Charlotte Islands, with consistently glabrous ovaries, was named subsp. glabellicarpa. Some southeastern Alaska populations have plants with glabrous, partially hairy, and completely short-silky ovaries growing together. The possibility that subsp. glabellicarpa may be a hybrid or a simple mutation needs study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Occurrence of Salix candida in Nunavut is on Akimiski Island in James Bay.

Salix candida is geographically wide-ranging but limited to calcareous habitats and, for that reason, it is quite local or even rare in some parts of its range.

Hybrids:

Salix candida forms natural hybrids with S. bebbiana, S. brachycarpa var. brachycarpa, S. calcicola, S. eriocephala, S. famelica, S. myrtillifolia, S. petiolaris, and S. planifolia. Hybrids with S. discolor, S. petiolaris, and S. sericea have been reported (the latter also by C. K. Schneider 1921; M. L. Fernald 1950) but no convincing specimens have been seen. Salix candida hybrids are recognized from their woolly indumentum that often is conspicuous on leaves, stems, and ovaries. In hybrids, these characters, especially woolly patches on ovaries, stand out as discordant variation.

Salix candida × S. eriocephala (S. ×rubella Bebb ex C. K. Schneider) was described by W. W. Rowlee and K. M. Wiegand (1896) as S. candida × S. cordata. In addition to woolly patches on the ovaries, they noted that buds of the hybrids usually are shorter, more divergent, and blunter than those in S. eriocephala, and are glabrous or hairy. Known from New York and Newfoundland; it should be expected throughout the sympatric range of the parental species.

Salix candida × S. famelica: The Saskatchewan specimen resembles S. famelica but has the leaf indumentum of S. candida.

Salix candida × S. myrtillifolia: Saskatchewan specimens combine characters of the two parents.

Salix candida × S. petiolaris: Intermediates between these species are known from Michigan and New York (W. W. Rowlee and K. M. Wiegand 1896) as well as Ontario and Saskatchewan, but can be expected wherever the two grow together. The invalid name “S. ×clarkei” is sometimes used for this hybrid.

The glabrescent form of Salix candida, forma denudata (Andersson) Rouleau, may be of hybrid origin.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 65. FNA vol. 7, p. 141.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia Salicaceae > Salix > subg. Vetrix > sect. Candidae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. reticulata var. gigantifolia, S. reticulata subsp. glabellicarpa, S. reticulata var. semicalva S. candida var. denudata
Name authority Linnaeus: Sp. Pl. 2: 1018. (1753) Flüggé ex Willdenow: Sp. Pl. 4: 708. (1806)
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