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bay-leaf or bay or laurel willow, bay-leaf willow, laurel willow

gray or small pussy or upland willow, prairie willow

Habit Shrubs or trees, 5–15 m. Stems: branches flexible at base, brownish or yellow-green, highly glossy, glabrous; branchlets yellow-green, red-brown, or brownish, glossy, glabrous. Shrubs, 0.3–3 m, (forming clones by layering).
Stems

branches dark red-brown, not or weakly glaucous, tomentose to glabrescent;

branchlets red-brown, yellow-brown, or greenish brown, moderately to very densely villous, tomentose, or velvety-tomentose to glabrescent.

Leaves

stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex rounded;

petiole deeply to shallowly grooved adaxially, 5–15 mm, with pairs or clusters of spherical glands distally or throughout, glabrous adaxially;

largest medial blade hypostomatous, narrowly elliptic, elliptic, or lanceolate, 50–135 × 20–50 mm, 2–4 times as long as wide, base convex, margins slightly revolute or flat, serrulate, apex acuminate, abaxial surface pale not glaucous, glabrous, adaxial highly glossy, glabrous;

proximal blade margins entire or serrulate;

juvenile blade reddish, glabrous abaxially.

stipules absent or rudimentary on early ones, absent, rudimentary or foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 0.5–7(–12) mm, velvety, pilose, or villous adaxially;

largest medial blade (sometimes hemiamphistomatous), narrowly oblong, narrowly elliptic, elliptic, oblanceolate, obovate, or broadly obovate, (13–)20–90(–135) × 3–23(–35) mm, 2.3–9 times as long as wide, base cuneate or convex, margins revolute or flat, entire, crenate, or sinuate, (glands submarginal), apex acuminate or convex, abaxial surface glaucous, sparsely to densely tomentose or woolly, hairs erect or spreading, wavy, adaxial slightly or highly glossy, glabrous, pubescent, tomentose, or pilose;

proximal blade margins entire or serrulate;

juvenile blade green, densely tomentose to glabrescent abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

abaxial nectary 0.6–1.7 mm, adaxial nectary square, ovate, or oblong, 0.5–1.5 mm, nectaries distinct or ± connate and cup-shaped;

stamens 4–10;

filaments distinct, hairy on proximal 1/2;

anthers ellipsoid or globose, 0.5–0.6 mm.

adaxial nectary oblong or square, 0.2–0.7 mm;

filaments distinct, glabrous or hairy basally;

anthers purple turning yellow, ellipsoid or cylindrical, 0.4–0.6 mm.

Pistillate flowers

(abaxial nectary present or absent), adaxial nectary oblong, square, or ovate, 0.4–0.8 mm, (nectaries distinct or connate and shallowly cup-shaped), shorter than or equal to stipe;

stipe 0.5–1.6 mm;

ovary pyriform, beak bulged below or tapering to styles;

ovules 18–22 per ovary;

styles connate or distinct, 0.4–0.6 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.4–0.6 mm.

adaxial nectary square, 0.4–0.8 mm, shorter than stipe;

stipe 1–2.5 mm;

ovary obclavate or pyriform, moderately densely to sparsely short-silky-villous (hairs refractive), beak slightly bulged below styles, (valves recurving in fruit);

ovules 6–12 per ovary;

styles (sometimes slightly distinct distally), 0.2–0.4 mm;

stigmas slenderly to broadly cylindrical, 0.2–0.56 mm.

Capsules

6–9 mm.

5–12 mm.

Catkins

staminate 27–81 × 9–13 mm, flowering branchlet 9–21 mm; pistillate moderately to densely flowered, slender or stout, 29–68 × 7–15 mm, flowering branchlet 9–42 mm;

floral bract 2–4 mm, apex acute or rounded to truncate, entire or toothed, abaxially sparsely hairy (mainly proximally), hairs wavy or straight.

flowering before leaves emerge; staminate 6.5–34 × 5–19 mm, flowering branchlet 0–1 mm; pistillate (and staminate) moderately to very densely flowered, stout, subglobose, or globose, 9–47(–55 in fruit) × 5.5–19 mm, flowering branchlet 0–4 mm;

floral bract brown, black, or bicolor, 0.8–2 mm, apex rounded or acute, abaxially moderately densely hairy, hairs (white), straight or wavy.

2n

= 76.

Salix pentandra

Salix humilis

Phenology Flowering late May-mid Jun.
Habitat Shores of streams and lakes, marshes, roadsides, waste places
Elevation 0-2300 m (0-7500 ft)
Distribution
from FNA
AK; CO; CT; DC; IA; IL; KY; MA; MD; ME; MN; MT; NC; NE; NH; NJ; NY; OH; PA; RI; SD; VA; VT; WI; WY; AB; BC; MB; NB; NF; NS; ON; QC; SK; Eurasia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; WI; WV; MB; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

The Ohio occurrence is based on information from T. Cooperrider (pers. comm.).

Only pistillate plants of Salix pentandra are known to occur in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Varieties humilis and tristis are almost sympatric. In Maine they occur in the same populations without evident intermediate forms, while in other places, e.g., Florida, Georgia, and Nebraska, they apparently intergrade. Usually, they are distinct in habit, leaf and catkin size, and, most conspicuously, the presence of foliaceous stipules in var. humilis and their absence in var. tristis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stipules on late leaves foliaceous; largest medial blades (20-)50-90(-135) mm; petioles (1.5-)3-7(-12) mm; staminate catkins 14.5-34 mm; pistillate catkins longer than 20 mm; peeled wood smooth or striate, striae dense, to 20 mm.
var. humilis
1. Stipules on late leaves absent or rudimentary; largest medial blades (13-)20-50(-70) mm; petioles 0.5-3(-6) mm; staminate catkins 6.5-13.5 mm; pistillate catkins shorter than 20 mm; peeled wood smooth or striate, striae sparse, to 2 mm
var. tristis
Source FNA vol. 7, p. 44. FNA vol. 7, p. 129.
Parent taxa Salicaceae > Salix > subg. Salix > sect. Salicaster Salicaceae > Salix > subg. Vetrix > sect. Cinerella
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Subordinate taxa
S. humilis var. humilis, S. humilis var. tristis
Name authority Linnaeus: Sp. Pl. 2: 1016. (1753) Marshall: Arbust. Amer., 140. (1785)
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