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gray-leaf Sierra willow, gray-leafed Sierra willow, Sierra willow

Habit Plants 0.5–2 m. Stems: branches dark red-brown or yellow-brown, not to strongly glaucous, glabrous; branchlets yellow-brown or red-brown, (not or weakly glaucous), pilose or pubescent, hairs straight, wavy, or geniculate, (inner membranaceous bud-scale layer free, separating from outer layer).
Leaves

stipules foliaceous, rudimentary, or absent on early ones, foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 4–9 mm, pilose adaxially;

largest medial blade (sometimes amphistomatous), lorate, narrowly oblong, narrowly elliptic, or oblanceolate, 35–95 × 7.5–20 mm, 3.4–7.1 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex acute, acuminate, or convex, abaxial surface glaucous (sometimes obscured by hairs), sparsely to moderately densely long- to short-silky or pubescent, hairs (white, sometimes also ferruginous), straight or wavy, adaxial dull or slightly glossy, sparsely or moderately densely pubescent or long- to short-silky, (hairs white, sometimes also ferruginous);

proximal blade margins entire or serrulate;

juvenile blade densely long-silky abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

adaxial nectary oblong or ovate, 0.6–1.1 mm;

filaments distinct or connate less than 1/2 their lengths, hairy basally;

anthers purple turning yellow, 0.6–1 mm.

Pistillate flowers

adaxial nectary oblong or flask-shaped, 0.7–1.3 mm, shorter than or equal to stipe;

stipe 0.8–2 mm;

ovary obclavate or pyriform, short-silky-villous, beak gradually tapering to or slightly bulged below styles;

ovules 15–16 per ovary;

styles 0.6–1 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or broadly cylindrical, 0.24–0.32–0.44 mm.

Capsules

5–10 mm.

Catkins

flowering as leaves emerge; staminate stout, 15.5–34 × 7–14 mm, flowering branchlet 1–8 mm; pistillate moderately densely flowered, stout, 20–55(–65 in fruit) × 11–13 mm, flowering branchlet 2–15 mm;

floral bract dark brown or bicolor, 1.2–2.5 mm, apex acute or rounded, abaxially hairy, hairs straight or wavy.

Salix

×jesupii Fernald: S. alba × S. lucida;

S. ×ehrhartiana of authors, not G.

Meyer

.

The

origin of this hybrid, between a European and a native species, is unknown.

It

is characterized by: shrubs or trees, 7–10 m;

branchlets red-brown;

petioles convex to shallowly grooved adaxially, 3–11 mm, with pairs or clusters of spherical or stalked glands distally;

largest medial leaf blade amphistomatous or hemiamphistomatous, abaxial surface glaucous, sparsely long-silky to glabrescent, hairs white and ferruginous;

floral bract apex acute to rounded, toothed or entire; pistillate flowering branchlet 8–16 mm, bract deciduous after flowering;

stamens 3–5;

stipes 0.3–0.6 mm;

ovary pyriform, glabrous;

ovules 12–14 per ovary;

styles 0.3–0.8 mm;

capsules 4–5 mm.

Salix orestera

Salix ×jesupii

Phenology Flowering late May-late Aug. Flowering late April–mid May.
Habitat Subalpine meadows, slopes, lakes, streams, granite substrates Edges of streams and lakes, in wet deciduous woods, sand dunes, and wet railroad rights-of-way.
Elevation 2100-4000 m (6900-13100 ft) 10–300m (0–1000ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
ON; PE; QC; SK [Ill., Ky., Mass., N.H., N.Y., Ohio, Pa., Vt., Va., Wash., W.Va., Wis. It is possibly introduced in Wash.]
[BONAP county map]
Discussion

Salix orestera is so similar to S. glauca var. villosa that it is sometimes included in S. glauca. It is phenetically most similar to S. eastwoodiae and S. lemmonii (G. W. Argus 1997), and the possibility that it is a hybrid or an alloploid involving those species needs study.

Salix orestera can be distinguished from S. glauca var. villosa in having largest medial blades usually narrower, 3.4–7.1 times as long as wide, abaxial surfaces distinctly silky with appressed hairs pointing toward apex, hairs sometimes ferruginous, secondary veins raised abaxially and adaxially, branches often strongly glaucous, and staminate flowers without abaxial nectaries; S. glauca var. villosa has largest medial blades usually broader, 2.2–3.9 times as long as wide, abaxial surfaces usually glabrescent or, if hairy, hairs unkempt, slightly spreading, hairs always white, secondary veins raised abaxially, flat or impressed adaxially, branches not glaucous, staminate flowers often with abaxial nectaries.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix ×jesupii was named by M. S. Bebb (1895) as a formula hybrid, S. alba × S. lucida, and Fernald based his binomial on Bebb’s exsiccatae. In North America, it was mistaken for the European S. ×ehrhartiana G. Meyer (S. alba Linnaeus × S. pentandra Linnaeus) (G. W. Argus 1986). Study of nectary morphology confirmed that the North American plants were not the same as the European hybrid. In herbaria, plants of S. ×jesupii often are misidentified as S. alba, S. euxina, S. ×fragilis, or S. lucida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 108. FNA vol. 7, p. 43.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Salix > sect. Salix
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. commutata var. rubicunda, S. glauca subsp. orestera, S. glauca var. orestera
Name authority C. K. Schneider: J. Arnold Arbor. 1: 164. (1920) Fernald
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