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gray-leaf Sierra willow, gray-leafed Sierra willow, Sierra willow

Tyrrell's willow

Habit Plants 0.5–2 m. Stems: branches dark red-brown or yellow-brown, not to strongly glaucous, glabrous; branchlets yellow-brown or red-brown, (not or weakly glaucous), pilose or pubescent, hairs straight, wavy, or geniculate, (inner membranaceous bud-scale layer free, separating from outer layer). Shrubs, 0.6–3.5 m, (sometimes forming clones by layering).
Stems

branches and branchlets red-brown, not to strongly glaucous, glabrous, (buds caprea-type or intermediate).

Leaves

stipules foliaceous, rudimentary, or absent on early ones, foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 4–9 mm, pilose adaxially;

largest medial blade (sometimes amphistomatous), lorate, narrowly oblong, narrowly elliptic, or oblanceolate, 35–95 × 7.5–20 mm, 3.4–7.1 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex acute, acuminate, or convex, abaxial surface glaucous (sometimes obscured by hairs), sparsely to moderately densely long- to short-silky or pubescent, hairs (white, sometimes also ferruginous), straight or wavy, adaxial dull or slightly glossy, sparsely or moderately densely pubescent or long- to short-silky, (hairs white, sometimes also ferruginous);

proximal blade margins entire or serrulate;

juvenile blade densely long-silky abaxially, hairs white, sometimes also ferruginous.

stipules rudimentary or foliaceous (often brownish) on early ones, foliaceous or rudimentary on late ones, (ca. 4 mm), apex acute;

petiole convex to flat, or shallowly grooved adaxially, 1–3.4–16 mm, glabrous, pilose, or short-silky adaxially;

largest medial blade (amphistomatous), narrowly elliptic, elliptic, oblanceolate, or obovate, 15–29–65 × 3.5–8.8–18 mm, 2.3–3.3–4.4 times as long as wide, base cuneate or convex, margins strongly to slightly revolute, entire, or very shallowly serrulate or shallowly serrulate-crenulate, apex acute, acuminate, or convex, abaxial surface glaucous, sparsely long-silky to glabrescent, hairs (ferruginous), straight, adaxial highly glossy, glabrous or sparsely short-silky (hairs white, sometimes also ferruginous);

proximal blade margins entire;

juvenile blade yellowish green or sometimes reddish, sparsely long-silky abaxially, hairs white and ferruginous.

Staminate flowers

adaxial nectary oblong or ovate, 0.6–1.1 mm;

filaments distinct or connate less than 1/2 their lengths, hairy basally;

anthers purple turning yellow, 0.6–1 mm.

adaxial nectary oblong, 0.8–1.1 mm;

filaments distinct, glabrous or hairy basally;

anthers purple turning yellow, ellipsoid, 0.4–0.7 mm.

Pistillate flowers

adaxial nectary oblong or flask-shaped, 0.7–1.3 mm, shorter than or equal to stipe;

stipe 0.8–2 mm;

ovary obclavate or pyriform, short-silky-villous, beak gradually tapering to or slightly bulged below styles;

ovules 15–16 per ovary;

styles 0.6–1 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or broadly cylindrical, 0.24–0.32–0.44 mm.

adaxial nectary oblong or flask-shaped, 0.6–1.1 mm, equal to or shorter than stipe;

stipe 0.2–1 mm (–1.4 mm in cultivation);

ovary pyriform, long-silky, beak gradually tapering to styles;

ovules 12–16 per ovary;

styles 0.6–1.2 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.44–0.55–0.75 mm.

Capsules

5–10 mm.

3.6–5 mm.

Catkins

flowering as leaves emerge; staminate stout, 15.5–34 × 7–14 mm, flowering branchlet 1–8 mm; pistillate moderately densely flowered, stout, 20–55(–65 in fruit) × 11–13 mm, flowering branchlet 2–15 mm;

floral bract dark brown or bicolor, 1.2–2.5 mm, apex acute or rounded, abaxially hairy, hairs straight or wavy.

flowering before leaves emerge; staminate stout, 14–35 × 12–16 mm, flowering branchlet 0 mm; pistillate densely flowered, stout, 17–51 × 10–13–22 mm, flowering branchlet 0–4 mm;

floral bract brown, black, or bicolor, 1–3.7 mm, apex acute to acuminate or rounded, abaxially hairy, hairs long, straight.

Salix orestera

Salix tyrrellii

Phenology Flowering late May-late Aug. Flowering mid Jun-mid Jul.
Habitat Subalpine meadows, slopes, lakes, streams, granite substrates Active sand dunes, shrubby tundra
Elevation 2100-4000 m (6900-13100 ft) 200-600 m (700-2000 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AB; NT; NU; SK
[BONAP county map]
Discussion

Salix orestera is so similar to S. glauca var. villosa that it is sometimes included in S. glauca. It is phenetically most similar to S. eastwoodiae and S. lemmonii (G. W. Argus 1997), and the possibility that it is a hybrid or an alloploid involving those species needs study.

Salix orestera can be distinguished from S. glauca var. villosa in having largest medial blades usually narrower, 3.4–7.1 times as long as wide, abaxial surfaces distinctly silky with appressed hairs pointing toward apex, hairs sometimes ferruginous, secondary veins raised abaxially and adaxially, branches often strongly glaucous, and staminate flowers without abaxial nectaries; S. glauca var. villosa has largest medial blades usually broader, 2.2–3.9 times as long as wide, abaxial surfaces usually glabrescent or, if hairy, hairs unkempt, slightly spreading, hairs always white, secondary veins raised abaxially, flat or impressed adaxially, branches not glaucous, staminate flowers often with abaxial nectaries.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix tyrrellii, first described from Lake Athabasca sand dunes in northwestern Saskatchewan and adjacent Alberta, is characterized by slender, amphistomatous leaves and relatively long, slender branchlets (G. W. Argus and J. W. Steele 1979). It probably evolved, in the past 10,000 years, from the widespread boreal S. planifolia. Originally, it was thought to be endemic to Lake Athabasca but recent collections from Nunavut and the Northwest Territories suggest that it may have a much wider range. Plants from Nunavut and the Northwest Territories do not have the long, slender branchlets found in sand dune populations, and identification is primarily based on the presence of amphistomatous leaves. This character, however, may be unreliable. For example, S. planifolia from 2610 m in the Glass Mountains, California, have amphistomatous leaves, suggesting that this character may be under environmental influence. The appropriate rank for this taxon remains uncertain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 108. FNA vol. 7, p. 140.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Vetrix > sect. Phylicifoliae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. commutata var. rubicunda, S. glauca subsp. orestera, S. glauca var. orestera S. planifolia subsp. tyrrellii
Name authority C. K. Schneider: J. Arnold Arbor. 1: 164. (1920) Raup: J. Arnold Arbor. 17: 231, plate 192. (1936)
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