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gray-leaf Sierra willow, gray-leafed Sierra willow, Sierra willow

false mountain willow, serviceberry willow

Habit Plants 0.5–2 m. Stems: branches dark red-brown or yellow-brown, not to strongly glaucous, glabrous; branchlets yellow-brown or red-brown, (not or weakly glaucous), pilose or pubescent, hairs straight, wavy, or geniculate, (inner membranaceous bud-scale layer free, separating from outer layer). Plants 1–6 m. Stems: branches red-brown or yellow-brown, not or weakly glaucous, (slightly or highly glossy), glabrous or glabrescent; branchlets yellow-green, red-brown, or brownish, glabrous, pilose, or densely villous, (inner membranaceous bud-scale layer free, separating from outer layer).
Leaves

stipules foliaceous, rudimentary, or absent on early ones, foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 4–9 mm, pilose adaxially;

largest medial blade (sometimes amphistomatous), lorate, narrowly oblong, narrowly elliptic, or oblanceolate, 35–95 × 7.5–20 mm, 3.4–7.1 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex acute, acuminate, or convex, abaxial surface glaucous (sometimes obscured by hairs), sparsely to moderately densely long- to short-silky or pubescent, hairs (white, sometimes also ferruginous), straight or wavy, adaxial dull or slightly glossy, sparsely or moderately densely pubescent or long- to short-silky, (hairs white, sometimes also ferruginous);

proximal blade margins entire or serrulate;

juvenile blade densely long-silky abaxially, hairs white, sometimes also ferruginous.

stipules foliaceous, (prominent, 5–9–15 mm), apex rounded to acute, sometimes acuminate;

petiole (usually reddish), shallowly grooved, or convex to flat adaxially, 6–20 mm, short-silky or velvety adaxially;

largest medial blade broadly to narrowly elliptic, or ovate to broadly obovate, (25–)30–86(–118) × 12–51 mm, 1.4–3 times as long as wide, base convex, rounded, cuneate, subcordate, cordate, or sometimes irregularly lobed, margins flat, serrulate or crenate, apex acute, acuminate, or convex, abaxial surface glaucous, glabrous, pubescent, or pilose, hairs wavy, adaxial slightly glossy or dull, glabrous, puberulent, pubescent, or pilose, midrib hairy;

proximal blade margins entire or serrulate;

juvenile blade reddish, glabrous or pubescent abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

adaxial nectary oblong or ovate, 0.6–1.1 mm;

filaments distinct or connate less than 1/2 their lengths, hairy basally;

anthers purple turning yellow, 0.6–1 mm.

adaxial nectary oblong, 0.3–1 mm;

filaments distinct or connate less than 1/2 their lengths, glabrous;

anthers purple turning yellow, 0.4–0.5 mm.

Pistillate flowers

adaxial nectary oblong or flask-shaped, 0.7–1.3 mm, shorter than or equal to stipe;

stipe 0.8–2 mm;

ovary obclavate or pyriform, short-silky-villous, beak gradually tapering to or slightly bulged below styles;

ovules 15–16 per ovary;

styles 0.6–1 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or broadly cylindrical, 0.24–0.32–0.44 mm.

adaxial nectary oblong, flask-shaped, 0.3–0.8 mm, shorter than stipe;

stipe 0.5–0.8–3 mm;

ovary pyriform or obclavate, glabrous, beak gradually tapering to styles;

ovules 18 per ovary;

styles (0.5–)0.7–1.8 mm;

stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.1–0.21–0.29 mm.

Capsules

5–10 mm.

4–7 mm.

Catkins

flowering as leaves emerge; staminate stout, 15.5–34 × 7–14 mm, flowering branchlet 1–8 mm; pistillate moderately densely flowered, stout, 20–55(–65 in fruit) × 11–13 mm, flowering branchlet 2–15 mm;

floral bract dark brown or bicolor, 1.2–2.5 mm, apex acute or rounded, abaxially hairy, hairs straight or wavy.

flowering before leaves emerge; staminate stout, 16–39 × 10–12 mm, flowering branchlet 0 mm; pistillate densely or moderately densely flowered, slender to globose, 17–73 × 8–20 mm, flowering branchlet 0–5 mm;

floral bract brown or black, 1–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight.

2n

= 38.

Salix orestera

Salix pseudomonticola

Phenology Flowering late May-late Aug. Flowering late Apr-early Jun.
Habitat Subalpine meadows, slopes, lakes, streams, granite substrates Moist fens in drainageways in white spruce forests, treed bogs, balsam poplar forests, floodplains
Elevation 2100-4000 m (6900-13100 ft) 0-2500 m (0-8200 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; ID; MN; MT; SD; WA; WY; AB; BC; MB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix orestera is so similar to S. glauca var. villosa that it is sometimes included in S. glauca. It is phenetically most similar to S. eastwoodiae and S. lemmonii (G. W. Argus 1997), and the possibility that it is a hybrid or an alloploid involving those species needs study.

Salix orestera can be distinguished from S. glauca var. villosa in having largest medial blades usually narrower, 3.4–7.1 times as long as wide, abaxial surfaces distinctly silky with appressed hairs pointing toward apex, hairs sometimes ferruginous, secondary veins raised abaxially and adaxially, branches often strongly glaucous, and staminate flowers without abaxial nectaries; S. glauca var. villosa has largest medial blades usually broader, 2.2–3.9 times as long as wide, abaxial surfaces usually glabrescent or, if hairy, hairs unkempt, slightly spreading, hairs always white, secondary veins raised abaxially, flat or impressed adaxially, branches not glaucous, staminate flowers often with abaxial nectaries.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix pseudomonticola is characterized by precocious flowering; catkins sessile; juvenile leaf blades, petioles, and proximal midribs reddish; stipules prominent; and leaves and branchlets sparsely hairy. Branches older than two years have a distinctive pattern, which consists of a series of longitudinal splits in epidermis produced as the branch expands. The edge of epidermis around the split, where it has separated from the branch, is yellow and contrasts with the red-brown branch to which the epidermis still adheres.

Vegetative specimens of Salix pseudomonticola with yellow-brown branches can be confused with S. famelica. They may be separated by their juvenile leaf margins prominently and closely gland-dotted; stipules usually prominent, sometimes early deciduous; leaves broader (1.4–3 times as long as wide versus 2.6–7 in S. famelica); and petioles slender and often longer in relation to blade length. The possibility of hybridization needs study.

Vegetative specimens of Salix pseudomonticola can be distinguished from S. pyrifolia by juvenile leaves reddish and almost always with some ferruginous hairs, versus yellowish-green and glabrous or with white hairs, and mature leaves usually dull adaxially versus glossy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 108. FNA vol. 7, p. 117.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Vetrix > sect. Hastatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. commutata var. rubicunda, S. glauca subsp. orestera, S. glauca var. orestera S. barclayi var. pseudomonticola
Name authority C. K. Schneider: J. Arnold Arbor. 1: 164. (1920) C. R. Ball: Contr. U.S. Natl. Herb. 22: 321. (1921)
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