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gray-leaf Sierra willow, gray-leafed Sierra willow, Sierra willow

Florida willow

Habit Plants 0.5–2 m. Stems: branches dark red-brown or yellow-brown, not to strongly glaucous, glabrous; branchlets yellow-brown or red-brown, (not or weakly glaucous), pilose or pubescent, hairs straight, wavy, or geniculate, (inner membranaceous bud-scale layer free, separating from outer layer).
Stems

branches red-brown, pubescent to glabrescent;

branchlets yellow-brown or red-brown, sparsely velvety or pubescent.

Leaves

stipules foliaceous, rudimentary, or absent on early ones, foliaceous on late ones, apex acute;

petiole convex to flat, or shallowly grooved adaxially, 4–9 mm, pilose adaxially;

largest medial blade (sometimes amphistomatous), lorate, narrowly oblong, narrowly elliptic, or oblanceolate, 35–95 × 7.5–20 mm, 3.4–7.1 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex acute, acuminate, or convex, abaxial surface glaucous (sometimes obscured by hairs), sparsely to moderately densely long- to short-silky or pubescent, hairs (white, sometimes also ferruginous), straight or wavy, adaxial dull or slightly glossy, sparsely or moderately densely pubescent or long- to short-silky, (hairs white, sometimes also ferruginous);

proximal blade margins entire or serrulate;

juvenile blade densely long-silky abaxially, hairs white, sometimes also ferruginous.

stipules absent or rudimentary on early ones, foliaceous on late ones, apex rounded;

petiole (shallowly grooved adaxially), 13–20 mm, puberulent or velvety adaxially;

largest medial blade narrowly oblong, oblong, lanceolate, or narrowly ovate, 100–170 × 38–55 mm, 2.5–3(–4) times as long as wide, base rounded, convex, or subcordate (cordate on larger leaves), margins serrulate or spinulose-serrulate, apex acute, acuminate, or convex, abaxial surface sparsely tomentose, hairs straight, adaxial highly glossy, sparsely villous or pilose to glabrescent (midrib remaining villous), hairs white and ferruginous;

proximal blade margins entire;

juvenile blade sparsely pubescent to very densely villous or pilose abaxially, hairs white.

Staminate flowers

adaxial nectary oblong or ovate, 0.6–1.1 mm;

filaments distinct or connate less than 1/2 their lengths, hairy basally;

anthers purple turning yellow, 0.6–1 mm.

abaxial nectary 0.4–0.8 mm, adaxial nectary ovate, 0.5–1.1 mm, nectaries distinct or connate and cup-shaped;

stamens 3–7;

filaments hairy basally;

anthers 0.4–0.5 mm.

Pistillate flowers

adaxial nectary oblong or flask-shaped, 0.7–1.3 mm, shorter than or equal to stipe;

stipe 0.8–2 mm;

ovary obclavate or pyriform, short-silky-villous, beak gradually tapering to or slightly bulged below styles;

ovules 15–16 per ovary;

styles 0.6–1 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or broadly cylindrical, 0.24–0.32–0.44 mm.

adaxial nectary square, 0.5–0.9 mm;

stipe 3.2–5.6 mm;

ovary obclavate to ellipsoidal, (rarely puberulent), beak gradually tapering to styles;

ovules 4 per ovary;

styles 0.3–0.4 mm;

stigmas 0.16–0.17–0.2 mm.

Capsules

5–10 mm.

6–7 mm.

Catkins

flowering as leaves emerge; staminate stout, 15.5–34 × 7–14 mm, flowering branchlet 1–8 mm; pistillate moderately densely flowered, stout, 20–55(–65 in fruit) × 11–13 mm, flowering branchlet 2–15 mm;

floral bract dark brown or bicolor, 1.2–2.5 mm, apex acute or rounded, abaxially hairy, hairs straight or wavy.

staminate 29–72 × 12–15 mm, flowering branchlet 1–10 mm; pistillate 50–81 × 17–27 mm, flowering branchlet 5–30 mm;

floral bract (tawny, sometimes greenish), 2–3.6 mm, apex rounded, entire, abaxially sparsely hairy, hairs wavy; pistillate bract persistent after flowering.

2n

= 38.

Salix orestera

Salix floridana

Phenology Flowering late May-late Aug. Flowering mid Feb-early Apr.
Habitat Subalpine meadows, slopes, lakes, streams, granite substrates Swamps, marshy shores of streams in woodlands, calcareous areas, shade tolerant
Elevation 2100-4000 m (6900-13100 ft) 10-40 m (0-100 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA
[BONAP county map]
Discussion

Salix orestera is so similar to S. glauca var. villosa that it is sometimes included in S. glauca. It is phenetically most similar to S. eastwoodiae and S. lemmonii (G. W. Argus 1997), and the possibility that it is a hybrid or an alloploid involving those species needs study.

Salix orestera can be distinguished from S. glauca var. villosa in having largest medial blades usually narrower, 3.4–7.1 times as long as wide, abaxial surfaces distinctly silky with appressed hairs pointing toward apex, hairs sometimes ferruginous, secondary veins raised abaxially and adaxially, branches often strongly glaucous, and staminate flowers without abaxial nectaries; S. glauca var. villosa has largest medial blades usually broader, 2.2–3.9 times as long as wide, abaxial surfaces usually glabrescent or, if hairy, hairs unkempt, slightly spreading, hairs always white, secondary veins raised abaxially, flat or impressed adaxially, branches not glaucous, staminate flowers often with abaxial nectaries.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

The closest relatives of Salix floridana are in the Old World sect. Tetraspermae. A detailed discussion of the distribution, taxonomy, and relationships of this uncommon subtropical endemic was given by G. W. Argus (1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 108. FNA vol. 7, p. 32.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Protitea > sect. Floridanae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. commutata var. rubicunda, S. glauca subsp. orestera, S. glauca var. orestera
Name authority C. K. Schneider: J. Arnold Arbor. 1: 164. (1920) Chapman: Fl. South. U.S., 430. (1860)
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