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dark sandbar willow, dusky willow

blanket-leaf willow

Habit Shrubs, 0.8–4 m. Stems: branches gray-brown or red-brown, glabrous or hairy; branchlets gray-brown to dark red-brown, glabrous, puberulent, densely long-silky, or villous to glabrescent. Shrubs, 1–3 m, (forming clones by layering).
Stems

branches yellow-brown, gray-brown, or red-brown, not glaucous, villous in patches to glabrescent;

branchlets gray-brown or red-brown, very densely villous.

Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones (apex acuminate);

petiole 1.5–8 mm, glabrous adaxially;

largest medial blade lorate, narrowly oblong, narrowly elliptic, narrowly oblanceolate, or linear, 30–133 × 5–20 mm, 3.4–8–15 times as long as wide, base cuneate or convex, margins flat, spinulose-serrulate or entire, apex acute, acuminate, or convex, abaxial surface glaucous or not, pilose, villous, or long-silky to glabrescent, hairs appressed or spreading, wavy, adaxial slightly glossy, villous to glabrescent;

proximal blade margins entire or serrulate;

juvenile blade reddish or yellowish green, densely villous abaxially.

stipules foliaceous, apex acute to acuminate;

petiole convex to flat adaxially, 5–12 mm, villous or tomentose adaxially, (strongly ventricose around floral buds);

largest medial blade (apparently hypostomatous but surfaces obscured by hairs), narrowly oblong, narrowly elliptic, elliptic, or obovate, 36–84 × 19–40 mm, 1.8–3.6 times as long as wide, base convex or cuneate, margins slightly revolute, entire, apex convex or acuminate, abaxial surface very densely woolly-tomentose, hairs wavy, adaxial dull, moderately to very densely, villous-tomentose;

proximal blade margins entire;

juvenile blade color obscured by hairs, very densely tomentose-woolly abaxially, hairs white.

Staminate flowers

abaxial nectary 0.3–0.9 mm, adaxial nectary narrowly oblong, oblong, or flask-shaped, 0.4–1.2 mm, nectaries distinct;

filaments densely hairy on proximal 1/2;

anthers 0.55–0.7–0.9 mm.

adaxial nectary oblong to narrowly oblong, 0.6–1.5 mm;

filaments distinct;

anthers yellow, ellipsoid, 0.6–0.7 mm.

Pistillate flowers

adaxial nectary ovate, oblong, or flask-shaped, 0.4–1.1 mm, longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.7 mm;

ovary obclavate or pyriform, glabrous, beak abruptly tapering to styles;

ovules 13–22 per ovary;

styles 0–0.14–0.5 mm;

stigmas slenderly cylindrical or 2 plump lobes, 0.2–0.5 mm.

adaxial nectary oblong or obtriangular, 0.5–1.1 mm;

stipe 0–0.3 mm;

ovary pyriform, beak gradually tapering to styles;

ovules 12–14 per ovary;

styles 1.2–2.2 mm;

stigmas 0.52–0.75–1 mm.

Capsules

4–5 mm.

4–7 mm.

Catkins

staminate 18–48 × 5–13 mm, flowering branchlet 3–15 mm; pistillate moderately densely flowered, slender or stout, 22–58 × 4–9 mm, flowering branchlet 4–12 mm;

floral bract (sometimes brown), 1.3–2.8 mm, apex rounded (sometimes truncate), entire or erose, abaxially hairy mainly proximally, hairs wavy.

flowering before leaves emerge; staminate stout, 40–56 × 14–15 mm, flowering branchlet 0 mm; pistillate densely flowered, slender, 35–125(–130 in fruit) mm, flowering branchlet 0 mm;

floral bract brown or black, 2–3 mm, apex convex to rounded, abaxially hairy, hairs straight.

2n

= 38.

Salix melanopsis

Salix silicicola

Phenology Flowering early May-mid Jul. No data are available on flowering time in the wild; in cultivation flowering is early May.
Habitat Riparian, floodplains, stream banks, subalpine meadows, coarse-textured substrates, silt Active sand dunes
Elevation 600-3100 m (2000-10200 ft) 20-500 m (100-1600 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
NU; SK
[BONAP county map]
Discussion

Salix fluviatilis Nuttall, long used for a Columbia River endemic (see 22. S. columbiana), is a rejected name.

Hybrids:

Salix melanopsis forms natural hybrids with S. exigua var. exigua, S. sessilifolia, and S. sitchensis (R. D. Dorn 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Comparisons of genetic variation in Salix alaxensis var. alaxensis from British Columbia and S. silicicola from Lake Athabasca sand dunes, Saskatchewan, based on isozyme loci, fit predicted relationships between progenitor and derived taxon (B. G. Purdy and R. J. Bayer 1995). Allelic diversity of S. silicicola was a subset of that of S. alaxensis, there was less genetic variation in S. silicicola than in S. alaxensis, and interspecific genetic variation within the two species was similar and relatively very high. This suggested a recent origin for the derived S. silicicola.

Salix silicicola is a uniform population that differs from S. alaxensis in its very densely villous or tomentose leaves and branchlets. These characters seem to be an adaptation to reduce sand abrasion and water loss in a sand dune environment. It is unlikely that it would have evolved in situ but probably derived from a pre-adapted source such as the one represented by specimens of putative S. silicicola from Pelly Lake, Nunavut. The isozyme study did not include specimens from that population or of S. alaxensis from Northwest Territories from which S. silicicola is likely to have been derived. Occurrence of S. silicicola-like plants in northern continental Nunavut suggests that during the late Pleistocene, it had a wider range, which now is represented by two disjunct populations. The question of appropriate taxonomic rank for the derived taxon is still unresolved. Although S. silicicola is different from S. alaxensis in its general appearance, they are very similar genetically, and argument could be made for treating them as varieties (B. Boivin 1966b).

Hybrids:

Salix silicicola forms natural hybrids with S. brachycarpa var. psammophila.

of conservation concern

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 58. FNA vol. 7, p. 147.
Parent taxa Salicaceae > Salix > subg. Longifoliae Salicaceae > Salix > subg. Vetrix > sect. Villosae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. bolanderiana, S. exigua var. gracilipes, S. exigua subsp. melanopsis, S. exigua var. tenerrima, S. fluviatilis var. tenerrima, S. longifolia var. tenerrima, S. melanopsis var. bolanderiana, S. melanopsis var. gracilipes, S. melanopsis var. kronkheitii, S. melanopsis var. tenerrima, S. parksiana, S. sessilifolia var. vancouverensis, S. tenerrima S. alaxensis var. silicicola
Name authority Nuttall: N. Amer. Sylv. 1: 78, plate 21. (1842) Raup: J. Arnold Arbor. 17: 236, plate 194. (1936)
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