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dark sandbar willow, dusky willow

bilberry willow, blueberry willow, low blueberry willow

Habit Shrubs, 0.8–4 m. Stems: branches gray-brown or red-brown, glabrous or hairy; branchlets gray-brown to dark red-brown, glabrous, puberulent, densely long-silky, or villous to glabrescent. Plants 0.1–0.6(–1) m, (forming clones by layering).
Stems

(decumbent);

branches gray-brown, red-brown, or yellow-brown, not to strongly glaucous (dull or slightly glossy), pubescent;

branchlets gray-brown, red-brown, or yellow-brown, sparsely pubescent.

Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones (apex acuminate);

petiole 1.5–8 mm, glabrous adaxially;

largest medial blade lorate, narrowly oblong, narrowly elliptic, narrowly oblanceolate, or linear, 30–133 × 5–20 mm, 3.4–8–15 times as long as wide, base cuneate or convex, margins flat, spinulose-serrulate or entire, apex acute, acuminate, or convex, abaxial surface glaucous or not, pilose, villous, or long-silky to glabrescent, hairs appressed or spreading, wavy, adaxial slightly glossy, villous to glabrescent;

proximal blade margins entire or serrulate;

juvenile blade reddish or yellowish green, densely villous abaxially.

stipules rudimentary, foliaceous, or absent on early ones, foliaceous on late ones (0.2–1.8(–5) mm);

petiole deeply to shallowly grooved adaxially, 1.5–8 mm, glabrous or pubescent adaxially;

largest medial blade (sometimes amphistomatous), elliptic, narrowly elliptic, obovate, or broadly obovate, 17–74 × 8–30 mm, 1.2–4.5 times as long as wide, base cuneate, convex, or subcordate, margins flat, serrulate, crenulate, or sinuate, apex acute, convex, or acuminate, abaxial surface not glaucous, glabrous, adaxial slightly glossy, glabrous;

proximal blade margins crenate;

juvenile blade reddish or yellowish green, glabrous.

Staminate flowers

abaxial nectary 0.3–0.9 mm, adaxial nectary narrowly oblong, oblong, or flask-shaped, 0.4–1.2 mm, nectaries distinct;

filaments densely hairy on proximal 1/2;

anthers 0.55–0.7–0.9 mm.

adaxial nectary oblong, ovate, or square, 0.2–0.34–0.4 mm;

filaments distinct, glabrous;

anthers purple turning yellow, 0.3–0.6 mm.

Pistillate flowers

adaxial nectary ovate, oblong, or flask-shaped, 0.4–1.1 mm, longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.7 mm;

ovary obclavate or pyriform, glabrous, beak abruptly tapering to styles;

ovules 13–22 per ovary;

styles 0–0.14–0.5 mm;

stigmas slenderly cylindrical or 2 plump lobes, 0.2–0.5 mm.

adaxial nectary square, oblong, or ovate, 0.2–0.4 mm, shorter than stipe;

stipe 0.6–1.7 mm;

ovary pyriform, glabrous, beak gradually tapering to or slightly bulged below styles;

ovules (6–)10–14 per ovary;

styles connate or distinct 1/2 their lengths, 0.3–0.7 mm;

stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.16–0.23–0.32 mm.

Capsules

4–5 mm.

4–6 mm.

Catkins

staminate 18–48 × 5–13 mm, flowering branchlet 3–15 mm; pistillate moderately densely flowered, slender or stout, 22–58 × 4–9 mm, flowering branchlet 4–12 mm;

floral bract (sometimes brown), 1.3–2.8 mm, apex rounded (sometimes truncate), entire or erose, abaxially hairy mainly proximally, hairs wavy.

flowering as leaves emerge; staminate stout, 11.5–39 × 5–14 mm, flowering branchlet 0.5–6 mm; pistillate moderately densely flowered or densely flowered, slender or stout, 16–46(–50 in fruit) × 4–15 mm, flowering branchlet 1.5–12 mm;

floral bract brown, black, tawny, or bicolor, 0.4–1.1 mm, apex retuse or acute, abaxially hairy throughout or proximally, hairs curly or wavy.

2n

= 38.

Salix melanopsis

Salix myrtillifolia

Phenology Flowering early May-mid Jul. Flowering early May-late Jul.
Habitat Riparian, floodplains, stream banks, subalpine meadows, coarse-textured substrates, silt Treed bogs, fens, stream banks, subalpine spruce thickets, Pinus contorta woods, sand dunes, coal spoils
Elevation 600-3100 m (2000-10200 ft) 90-2800 m (300-9200 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; WY; AB; BC; MB; NB; NT; NU; ON; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix fluviatilis Nuttall, long used for a Columbia River endemic (see 22. S. columbiana), is a rejected name.

Hybrids:

Salix melanopsis forms natural hybrids with S. exigua var. exigua, S. sessilifolia, and S. sitchensis (R. D. Dorn 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix myrtillifolia occurs in Nunavut on Akimiski Island in James Bay.

The complex of species related to Salix myrtillifolia includes S. arizonica, S. ballii, S. boothii, and S. pseudomyrsinites. Two are diploid (S. arizonica and S. myrtillifolia), and two are tetraploid (S. boothii and S. pseudomyrsinites); the chromosome number of S. ballii is unknown. They have been treated taxonomically in different ways, but are relatively distinct in their morphology, ecology, and geography. Salix myrtillifolia has outlying populations represented by single collections each from Colorado, Quebec, and Wyoming. Specimens attributed to this species from the Gaspe Peninsula, Quebec, and the Northern Peninsula, Newfoundland, all have evidence of leaf glaucescence and are S. ballii. See 57. S. pseudomyrsinites and 58. S. ballii for more description.

Hybrids:

Salix myrtillifolia forms natural hybrids with S. candida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 58. FNA vol. 7, p. 109.
Parent taxa Salicaceae > Salix > subg. Longifoliae Salicaceae > Salix > subg. Vetrix > sect. Hastatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. bolanderiana, S. exigua var. gracilipes, S. exigua subsp. melanopsis, S. exigua var. tenerrima, S. fluviatilis var. tenerrima, S. longifolia var. tenerrima, S. melanopsis var. bolanderiana, S. melanopsis var. gracilipes, S. melanopsis var. kronkheitii, S. melanopsis var. tenerrima, S. parksiana, S. sessilifolia var. vancouverensis, S. tenerrima
Name authority Nuttall: N. Amer. Sylv. 1: 78, plate 21. (1842) Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 132. (1858)
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