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dark sandbar willow, dusky willow

Geyer willow, Geyer's willow

Habit Shrubs, 0.8–4 m. Stems: branches gray-brown or red-brown, glabrous or hairy; branchlets gray-brown to dark red-brown, glabrous, puberulent, densely long-silky, or villous to glabrescent. Plants 0.6–5 m, (sometimes forming clones by stem fragmentation).
Stems

branches (sometimes ± brittle at base), yellow-green, gray-brown, red-brown, or violet, usually glaucous, glabrous or sparsely tomentose;

branchlets yellowish, yellow-brown, red-brown, or violet, (strongly glaucous or not), glabrous or sparsely to moderately densely pubescent, (buds caprea-type).

Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones (apex acuminate);

petiole 1.5–8 mm, glabrous adaxially;

largest medial blade lorate, narrowly oblong, narrowly elliptic, narrowly oblanceolate, or linear, 30–133 × 5–20 mm, 3.4–8–15 times as long as wide, base cuneate or convex, margins flat, spinulose-serrulate or entire, apex acute, acuminate, or convex, abaxial surface glaucous or not, pilose, villous, or long-silky to glabrescent, hairs appressed or spreading, wavy, adaxial slightly glossy, villous to glabrescent;

proximal blade margins entire or serrulate;

juvenile blade reddish or yellowish green, densely villous abaxially.

stipules usually absent or rudimentary (rarely foliaceous);

petiole convex to flat, or shallowly to deeply grooved adaxially, 2–9 mm, velvety, short-silky, or pubescent adaxially;

largest medial blade lorate, narrowly elliptic, or linear, 32–89 × 5.5–14 mm, 3.6–8.4(–11.3) times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire or distantly and shallowly serrulate, apex acute to acuminate, abaxial surface glaucous, glabrous or densely short- or long-silky, hairs (white, sometimes also ferruginous), straight, adaxial slightly glossy, densely short- or long-silky, especially midrib, to glabrescent, (hairs white, sometimes also ferruginous);

proximal blade margins entire or distantly and shallowly serrulate;

juvenile blade reddish or yellowish green, densely to sparsely long- or short-silky abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

abaxial nectary 0.3–0.9 mm, adaxial nectary narrowly oblong, oblong, or flask-shaped, 0.4–1.2 mm, nectaries distinct;

filaments densely hairy on proximal 1/2;

anthers 0.55–0.7–0.9 mm.

adaxial nectary oblong, ovate, or square, 0.3–0.8(–0.9) mm;

filaments distinct, glabrous or hairy on proximal 1/2 or basally;

anthers yellow or purple turning yellow, ellipsoid or globose, 0.4–0.5(–0.6) mm.

Pistillate flowers

adaxial nectary ovate, oblong, or flask-shaped, 0.4–1.1 mm, longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.7 mm;

ovary obclavate or pyriform, glabrous, beak abruptly tapering to styles;

ovules 13–22 per ovary;

styles 0–0.14–0.5 mm;

stigmas slenderly cylindrical or 2 plump lobes, 0.2–0.5 mm.

adaxial nectary oblong, square, or ovate, 0.2–1 mm;

stipe (0.4–)1–2.8 mm;

ovary pyriform (gourd-shaped), beak gradually tapering to styles;

ovules 6–12 per ovary;

styles 0.1–0.2(–0.6) mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.2–0.3–0.44 mm.

Capsules

4–5 mm.

(3–)4–6 mm.

Catkins

staminate 18–48 × 5–13 mm, flowering branchlet 3–15 mm; pistillate moderately densely flowered, slender or stout, 22–58 × 4–9 mm, flowering branchlet 4–12 mm;

floral bract (sometimes brown), 1.3–2.8 mm, apex rounded (sometimes truncate), entire or erose, abaxially hairy mainly proximally, hairs wavy.

flowering (before or) as leaves emerge; staminate globose, (1–1.1–)11–18 × 6–11 mm, flowering branchlet 1–5 mm; pistillate densely to loosely flowered, subglobose to globose, 8–21 × 7–17 mm, flowering branchlet 0.5–8 mm;

floral bract tawny or brown (black), 1.2–2.8 mm, apex rounded or acute, abaxially hairy, hairs short, wavy or straight.

2n

= 38.

Salix melanopsis

Salix geyeriana

Phenology Flowering early May-mid Jul. Flowering late Apr-late Jun.
Habitat Riparian, floodplains, stream banks, subalpine meadows, coarse-textured substrates, silt Lowland wet streamsides, lakeshores, sedge meadows, springs, seepages, swamps, cienegas, fine-textured substrates
Elevation 600-3100 m (2000-10200 ft) 10-3300 m (0-10800 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix fluviatilis Nuttall, long used for a Columbia River endemic (see 22. S. columbiana), is a rejected name.

Hybrids:

Salix melanopsis forms natural hybrids with S. exigua var. exigua, S. sessilifolia, and S. sitchensis (R. D. Dorn 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix geyeriana is characterized by its dark gray appearance, slender, dark branches, narrow leaves long-silky on both surfaces, general absence of stipules, and small, subglobose catkins. Plants in the Pacific Northwest with foliaceous stipules may be hybrids or introgressants, but the other parent is unknown.

Hybrids:

Salix geyeriana forms natural hybrids with S. bebbiana, S. irrorata, S. lemmonii, S. ligulifolia, and S. pedicellaris. Alleged hybrids with S. sitchensis, based on plants from British Columbia with broader, more hairy leaves, and catkins longer than in S. geyeriana, but with the short stipes of S. sitchensis (J. K. Henry 1915), are unconvincing.

Salix geyeriana × S. irrorata: A series of specimens from Arizona usually resemble S. geyeriana but have some characters of S. irrorata: foliaceous stipules, toothed leaf margins, catkins flowering before leaves emerge, shorter stipes (0.4–1.2 mm), and longer styles (0.2–0.6 mm). They also have some unique characters: proximal leaves sometimes serrulate, leaves sometimes amphistomatous, adaxial leaf surfaces mostly with ferruginous hairs, and ovaries sometimes gourd-shaped.

Salix geyeriana × S. lemmonii is uncommon but in mixed stands of the parental species some plants resemble S. geyeriana in having relatively short, subspherical catkins, small anthers, and petioles sometimes with petiolar glands; and S. lemmonii in having leaf blades amphistomatous, margins serrulate, and foliaceous stipules on early leaves. Because the species have different chromosome numbers, hybrids may be infertile, but occasional seeds have been seen. This hybrid is known from California (Lassen and Sierra counties), Oregon (Jefferson and Lane counties), and near Victoria, British Columbia.

Salix geyeriana × S. ligulifolia: Plants in Arizona with rudimentary stipules and leaves with ferruginous hairs may be this hybrid.

Salix geyeriana × S. pedicellaris occurs in Washington. It has the white and ferruginous hairs on leaves and ovaries of S. geyeriana, and leaves glaucous adaxially with prominent 2 and 3 veins of S. pedicellaris.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 58. FNA vol. 7, p. 153.
Parent taxa Salicaceae > Salix > subg. Longifoliae Salicaceae > Salix > subg. Vetrix > sect. Geyerianae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. bolanderiana, S. exigua var. gracilipes, S. exigua subsp. melanopsis, S. exigua var. tenerrima, S. fluviatilis var. tenerrima, S. longifolia var. tenerrima, S. melanopsis var. bolanderiana, S. melanopsis var. gracilipes, S. melanopsis var. kronkheitii, S. melanopsis var. tenerrima, S. parksiana, S. sessilifolia var. vancouverensis, S. tenerrima S. geyeriana var. argentea, S. geyeriana var. meleina
Name authority Nuttall: N. Amer. Sylv. 1: 78, plate 21. (1842) Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 122. (1858)
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