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dark sandbar willow, dusky willow

Columbia River willow

Habit Shrubs, 0.8–4 m. Stems: branches gray-brown or red-brown, glabrous or hairy; branchlets gray-brown to dark red-brown, glabrous, puberulent, densely long-silky, or villous to glabrescent. Shrubs, 2–6.5 m. Stems: branches red-brown, glabrous or hairy at nodes; branchlets yellow-brown or red-brown, pubescent, puberulent, or glabrous, (hairs appressed, straight or wavy).
Leaves

stipules absent, rudimentary, or foliaceous on early ones, foliaceous on late ones (apex acuminate);

petiole 1.5–8 mm, glabrous adaxially;

largest medial blade lorate, narrowly oblong, narrowly elliptic, narrowly oblanceolate, or linear, 30–133 × 5–20 mm, 3.4–8–15 times as long as wide, base cuneate or convex, margins flat, spinulose-serrulate or entire, apex acute, acuminate, or convex, abaxial surface glaucous or not, pilose, villous, or long-silky to glabrescent, hairs appressed or spreading, wavy, adaxial slightly glossy, villous to glabrescent;

proximal blade margins entire or serrulate;

juvenile blade reddish or yellowish green, densely villous abaxially.

stipules usually absent or rudimentary, sometimes foliaceous on early ones (apex acute), rudimentary or foliaceous on late ones;

petiole 2–5 mm, short-silky adaxially;

largest medial blade (sometimes hypostomatous, glands marginal), linear or very narrowly elliptic, 58–115 × 5–17 mm, 5.7–12.8 times as long as wide, base acute or cuneate, margins flat to slightly revolute, remotely spinulose-serrulate or sinuate, apex acuminate, abaxial surface glaucous (sometimes obscured by hairs), sparsely to very densely short- or long-silky, hairs appressed to slightly spreading, straight, adaxial dull, pubescent or long-silky;

proximal blade margins entire or remotely denticulate;

juvenile blade color obscured by hairs, densely short-silky abaxially.

Staminate flowers

abaxial nectary 0.3–0.9 mm, adaxial nectary narrowly oblong, oblong, or flask-shaped, 0.4–1.2 mm, nectaries distinct;

filaments densely hairy on proximal 1/2;

anthers 0.55–0.7–0.9 mm.

abaxial nectary 0.3–0.9 mm, adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.5–1 mm, nectaries distinct or connate and shallowly cup-shaped;

filaments hairy on proximal 1/2, throughout, or basally;

anthers 0.8–1.3 mm.

Pistillate flowers

adaxial nectary ovate, oblong, or flask-shaped, 0.4–1.1 mm, longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0–0.7 mm;

ovary obclavate or pyriform, glabrous, beak abruptly tapering to styles;

ovules 13–22 per ovary;

styles 0–0.14–0.5 mm;

stigmas slenderly cylindrical or 2 plump lobes, 0.2–0.5 mm.

(abaxial nectary 0.3–0.4 mm), adaxial nectary narrowly oblong, ovate, or flask-shaped, 0.4–1.3 mm, longer than stipe, nectaries distinct or connate and shallowly or partially cup-shaped;

stipe 0.2–0.7 mm;

ovary pyriform, long-silky, beak abruptly tapering to styles;

ovules 18–30 per ovary;

styles 0.1–0.2(–0.4) mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, (0.3–)0.5–1.1 mm.

Capsules

4–5 mm.

3.4–5.6 mm.

Catkins

staminate 18–48 × 5–13 mm, flowering branchlet 3–15 mm; pistillate moderately densely flowered, slender or stout, 22–58 × 4–9 mm, flowering branchlet 4–12 mm;

floral bract (sometimes brown), 1.3–2.8 mm, apex rounded (sometimes truncate), entire or erose, abaxially hairy mainly proximally, hairs wavy.

staminate 20–83 × 5–13 mm, flowering branchlet 6–125(–200) mm; pistillate densely to moderately densely flowered, slender or stout, 35–90 × 5–12 mm, flowering branchlet 11–160 mm;

floral bract 1.4–4 mm, apex acute, rounded, retuse, or truncate, entire or toothed, abaxially hairy throughout or proximally, hairs straight or wavy.

2n

= 38.

Salix melanopsis

Salix columbiana

Phenology Flowering early May-mid Jul. Flowering May-late Jul.
Habitat Riparian, floodplains, stream banks, subalpine meadows, coarse-textured substrates, silt Riparian, sandy-silty to sandy gravel floodplains, old beach dunes, rocky fill along streams
Elevation 600-3100 m (2000-10200 ft) 5-40 m (0-100 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; WA
[BONAP county map]
Discussion

Salix fluviatilis Nuttall, long used for a Columbia River endemic (see 22. S. columbiana), is a rejected name.

Hybrids:

Salix melanopsis forms natural hybrids with S. exigua var. exigua, S. sessilifolia, and S. sitchensis (R. D. Dorn 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybrids:

Salix columbiana forms natural hybrids with S. exigua var. exigua and S. sessilifolia. Both hybrids are reported from Oregon and Washington (R. D. Dorn 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 58. FNA vol. 7, p. 59.
Parent taxa Salicaceae > Salix > subg. Longifoliae Salicaceae > Salix > subg. Longifoliae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. bolanderiana, S. exigua var. gracilipes, S. exigua subsp. melanopsis, S. exigua var. tenerrima, S. fluviatilis var. tenerrima, S. longifolia var. tenerrima, S. melanopsis var. bolanderiana, S. melanopsis var. gracilipes, S. melanopsis var. kronkheitii, S. melanopsis var. tenerrima, S. parksiana, S. sessilifolia var. vancouverensis, S. tenerrima S. exigua var. columbiana
Name authority Nuttall: N. Amer. Sylv. 1: 78, plate 21. (1842) (Dorn) Argus: Harvard Pap. Bot. 12: 359. (2007)
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