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Pacific willow (var. lasiandra), shining willow, whiplash willow (ssp. caudata)

Turnor's willow

Habit Shrubs or trees, 4–6 m. Stems: branches flexible to highly brittle at base, yellow-brown, gray-brown, or red-brown, slightly to highly glossy, glabrous or villous to glabrescent; branchlets yellow-brown, gray-brown, or red-brown, glabrous, pilose, densely villous, or velvety, hairs spreading, straight, wavy, or crinkled. Shrubs, 1–2.5 m, (forming clones by layering).
Stems

branches yellow-brown or yellow-gray, not or weakly glaucous, (with sparkling wax crystals, dull or slightly glossy), pilose or villous;

branchlets gray-brown or red-brown, pubescent, villous, or velvety.

Leaves

stipules foliaceous, apex convex to rounded;

petiole shallowly to deeply grooved adaxially, 5–13 mm, with clusters of spherical or foliaceous glands distally, glabrous, pilose, or densely villous adaxially;

largest medial blade usually hypostomatous or hemiamphistomatous, rarely amphistomatous, lorate, very narrowly elliptic, narrowly elliptic, or lanceolate, (24–)55–133 × 11–43 mm, 2.5–6.2 times as long as wide, base convex or cuneate, margins flat, serrulate, apex acuminate to caudate, abaxial surface usually not glaucous (rarely so), glabrous, pilose, or moderately densely villous or long-silky, hairs appressed or spreading, white and/or ferruginous, straight or wavy, (coarse, caducous), adaxial (secondary veins flat or protruding), slightly or highly glossy, glabrous, pilose, or long-silky, hairs white and/or ferruginous;

proximal blade margins entire and glandular-dotted, or serrulate or crenulate;

juvenile blade reddish or yellowish green, glabrous or densely villous or long-silky abaxially, hairs white and ferruginous.

stipules foliaceous, apex acute or convex;

petiole shallowly grooved adaxially, 4–13 mm, villous or pubescent adaxially;

largest medial blade (amphistomatous), narrowly oblong, narrowly elliptic, elliptic, oblanceolate, or lanceolate, 26–47 × 7.5–15 mm, 2.8–4.1 times as long as wide, base convex, rounded, or subcordate, margins slightly revolute or flat, serrate or serrulate, apex acuminate to acute, abaxial surface glaucous, glabrous, pilose, villous, or long-silky, hairs straight or wavy, adaxial dull, sparsely or moderately densely pilose or long-silky, especially on midrib;

proximal blade margins entire or serrulate;

juvenile blade reddish or yellowish green, sparsely to moderately densely long-silky or pubescent abaxially, hairs white.

Staminate flowers

abaxial nectary 0.5–1.1 mm, adaxial nectary square or ovate, 0.3–0.9 mm, nectaries connate and cup-shaped;

stamens 3–6;

filaments distinct, hairy on proximal 1/2 or basally;

anthers ellipsoid, shortly cylindrical, obovoid, or globose, 0.6–0.8 mm.

adaxial nectary narrowly oblong or ovate, 0.7–1.1 mm;

filaments distinct or connate, hairy basally;

anthers yellow, 0.6–0.8 mm.

Pistillate flowers

adaxial nectary square or ovate, (swollen), 0.2–0.5 mm, shorter than stipe;

stipe 0.5–2 mm;

ovary pyriform, beak slightly bulged below or gradually tapering to styles;

ovules 18–24 per ovary;

styles connate or distinct 1/2 their lengths, 0.5–0.8 mm;

stigmas flat, abaxially non-papillate with rounded tip, broadly cylindrical, or 2 plump lobes, 0.24–0.31–0.42 mm.

adaxial nectary oblong, flask-shaped, or narrowly ovate, 0.4–1 mm, shorter than stipe;

stipe 2–4 mm;

ovary pyriform, glabrous, beaks slightly bulged below styles;

ovules 14–18 per ovary;

styles 0.3–0.5 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.16–0.23–0.28 mm.

Capsules

5–7 mm.

2.5–5 mm.

Catkins

staminate 19–69 × 4–14 mm, flowering branchlet 5–23 mm; pistillate (fruiting in summer), moderately densely to loosely flowered, slender to stout, 23–56(–70 in fruit) × 8–12 mm, flowering branchlet 8–25 mm;

floral bract 1.5–3 mm, apex convex or rounded, entire or toothed, abaxially sparsely hairy throughout or proximally, hairs wavy.

flowering just before or as leaves emerge; staminate stout, 16–30 × 8–10 mm, flowering branchlet 1.5–4 mm; pistillate loosely flowered, stout, 18–22 × 9–11 mm, flowering branchlet 3–4 mm;

floral bract brown or tawny, 1.2–1.6 mm, apex acute or rounded, abaxially hairy throughout or proximally, hairs straight or wavy.

2n

= 76.

Salix lucida

Salix turnorii

Phenology Flowering early May-mid Jul. No flowering time data are available (probably May or Jun).
Habitat Sandy or gravelly floodplains, lake margins, sedge meadows, vernal pools, alvars, open fens, marl bogs, treed bogs Active sand dunes
Elevation 0-600 m (0-2000 ft) 200-300 m (700-1000 ft)
Distribution
from FNA
CT; DE; IA; IL; IN; MA; MD; ME; MI; MN; ND; NH; NJ; NY; OH; PA; RI; SD; VA; VT; WI; WV; MB; NB; NL; NS; ON; PE; QC; SK; SPM
[WildflowerSearch map]
[BONAP county map]
from FNA
SK
[BONAP county map]
Discussion

The Virginia plants of Salix lucida are introduced (G. W. Argus 1986).

The Salix lucida complex is a group of three weakly delimited taxa, S. lasiandra var. caudata, S. lasiandra var. lasiandra, and S. lucida. The morphological characters used to separate them (leaves amphistomatous or hypostomatous and blades glaucous abaxially or not) are usually geographically correlated, but there are exceptions. G. W. Argus (1986b) proposed, based on principal components analysis of morphological data, to treat them as a single species consisting of three subspecies. The geographic overlap of the northeastern S. lucida and the western S. lasiandra is a relatively small area in central Saskatchewan. Evidence of intergradation was based on cultivation of a plant that, in the wild, had leaves that were not glaucous abaxially but were glaucous in cultivation. It seems best to treat them as two species, S. lucida and S. lasiandra, the latter with two varieties, var. lasiandra and var. caudata.

Hybrids:

Salix lucida forms natural hybrids with S. alba and S. nigra. Hybrids with S. serissima have been reported (M. L. Fernald 1950) but no convincing specimens have been seen. Attempts to hybridize S. lucida with members of subg. Protitea (S. amygdaloides), subg. Longifoliae (S. interior), and subg. Vetrix (S. discolor, S. eriocephala, and S. petiolaris) were unsuccessful (A. Mosseler 1990).

Salix lucida × S. nigra (S. ×schneideri B. Boivin) seems to be a rare intersubgeneric hybrid between tetraploid S. lucida and diploid S. nigra. It is known only from the type specimen, an infertile plant, growing with both parents. It resembles S. lucida in bud-scale margins connate, in petiolar glands stalked or foliaceous, and in leaf shape, and S. nigra in stipules rudimentary on proximal leaves and sometimes even on early leaves, stipule apex acute, pistillate catkins relatively long and slender, and styles relatively short.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Salix turnorii is known from the Lake Athabasca sand dunes in northwestern Saskatchewan. Salix famelica in the Great Sand Hills, southern Saskatchewan, is very similar morphologically and may have been the source of populations ancestral to S. turnorii that moved into northern Saskatchewan during the warm Holocene Hypsithermal Period (ca. 9000–6000 yrs. B.P.).

Hybrids:

Salix turnorii forms natural hybrids with S. brachycarpa var. psammophila.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 46. FNA vol. 7, p. 122.
Parent taxa Salicaceae > Salix > subg. Salix > sect. Salicaster Salicaceae > Salix > subg. Vetrix > sect. Cordatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms Pleiarina lucida, S. lucida var. angustifolia, S. lucida var. intonsa S. lutea var. turnorii
Name authority Muhlenberg: Ges. Naturf. Freunde Berlin Neue Schriften 4: 239, plate 6, fig. 7. (1803) Raup: J. Arnold Arbor. 17: 234, plate 193. (1936)
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