Salix humboldtiana |
Salix sphenophylla |
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Humboldt's willow |
wedge-leaf willow |
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Habit | Plants 0.03–0.12 m, not clonal or forming clones by layering. | |
Stems | trailing and rooting; branches yellow-brown or brownish, glabrous; branchlets yellow-brown, glabrous. |
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Leaves | stipules absent or rudimentary; petiole 4–25 mm, (glabrous or pilose adaxially); largest medial blade hypostomatous, narrowly elliptic, broadly elliptic, obovate, or very broadly obovate, 19–52 × 10–28 mm, 1–3 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex convex, retuse, or rounded, abaxial surface glabrous, pilose or sparsely long-silky to glabrescent, hairs straight or wavy, adaxial slightly glossy, glabrous or pilose; proximal blade margins entire; juvenile blade (reddish), very sparsely long-silky abaxially. |
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Staminate flowers | abaxial nectary absent, adaxial nectary oblong, 0.6–1 mm; filaments distinct; anthers ellipsoid or shortly cylindrical, 0.4–0.6 mm. |
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Pistillate flowers | adaxial nectary oblong or ovate, 0.7–1.6 mm, equal to or longer than stipe; stipe 0.5–1.4 mm; ovary obclavate or pyriform, glabrous, or patchy or streaky pilose or villous, especially on beak, beak gradually tapering to or slightly bulged below styles; ovules 10–18 per ovary; styles 0.6–1.8 mm; stigmas flat, abaxially non-papillate with pointed tip, or slenderly cylindrical, 0.32–0.5–0.68 mm. |
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Capsules | 4–12 mm. |
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Salix | humboldtiana Willdenow: Humboldt willow is not known to occur in the flora area. |
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It | is characterized by: trees, 4–25 m; branches highly brittle at base, bud-scale margins distinct and overlapping adaxially; stipules on late leaves rudimentary or foliaceous; largest medial leaf blade usually linear, abaxial surface not glaucous, adaxial dull; pistillate bract deciduous after flowering; stamens 3–7; capsules with distinct, often raised, white veins.; it occurs throughout much of Mexico to central Chile. |
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Catkins | staminate 21–53 × 7–13 mm, flowering branchlet 8–20 mm; pistillate loosely to densely flowered, slender or stout, 32–79 × 7–18 mm, flowering branchlet 4–27 mm; floral bract brown or black, 1.1–2 mm, apex rounded, entire, abaxially hairy or ciliate, hairs straight. |
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2n | = 38, 57. |
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Salix humboldtiana |
Salix sphenophylla |
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Phenology | Flowering mid Jun-late Jul (early Aug). | |
Habitat | Stony or gravelly substrates on talus, rocky outcrops, dry, stony tundra, sandy and moss tundra | |
Elevation | 10-900 m (0-3000 ft) | |
Distribution |
Mexico to central Chile |
AK; NT; YT; e Asia (Chukotka, Russian Far East, e Siberia) |
Discussion | Salix humboldtiana is closely related to S. nigra in its generally narrow leaf blades, which are not glaucous abaxially. The two differ in the following characters: S. humboldtiana has leaf blades linear to sometimes narrowly oblong (10–28.6 times as long as wide), ovaries usually ovoid to ellipsoid, ovary walls often stomatiferous and with raised, white veins, and capsule valves relatively thick, slightly recurved. S. nigra has leaf blades usually narrowly lanceolate (6–13 times as long as wide), ovaries pyriform to obclavate, ovary walls neither stomatiferous nor notably veined, and capsule valves relatively thin and strongly recurved. Both species occur in Chihuahua, Mexico. The report by R. I. Lonard et al. (1991) that specimens identified as Salix nigra from the lower Rio Grande, Texas, resemble S. humboldtiana in having strongly veined capsules suggests that S. humboldtiana, or intergrades with that species, may occur in Texas. Attempts to locate a voucher specimen were unsuccessful; because strongly veined capsules are diagnostic, further field study is indicated. An earlier name, Salix chilensis Molina, has been applied to this species; it does not seem to pertain to this taxon (C. K. Schneider 1918). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The patchy or streaky indumentum on the ovaries of some plants suggests that they may be hybrids. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 34. | FNA vol. 7, p. 82. |
Parent taxa | Salicaceae > Salix > subg. Protitea > sect. Humboldtianae | Salicaceae > Salix > subg. Chamaetia > sect. Diplodictyae |
Sibling taxa | ||
Synonyms | S. sphenophylla subsp. pseudotorulosa | |
Name authority | Willdenow | A. K. Skvortsov: Spisok Rast. Gerb. Fl. S.S.S.R. Bot. Inst. Vsesoyuzn. Akad. Nauk 16: 62. (1966) |
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