Salix humboldtiana
Salix pedicellaris
Humboldt's willow
bog willow
erect, decumbent or trailing;
branches gray-brown, glabrous;
branchlets yellow-brown or red-yellow, glabrous or puberulent (hairs straight, minute, inner membranaceous bud-scale layer free, not separating from outer layer).
stipules absent or rudimentary;
petiole (deeply to shallowly grooved adaxially), 3–8 mm, (glabrous or puberulent);
largest medial blade narrowly oblong, oblong, narrowly to broadly elliptic, narrowly oblanceolate or oblanceolate, 19–53(–69) × 5–20 mm, 1.8–4.9 times as long as wide, base convex or rounded, margins flat or slightly revolute, entire, apex acute, convex, or rounded, abaxial surface glabrous, adaxial dull, glaucous, glabrous (rarely very sparsely short-silky, hairs usually white, sometimes also ferruginous);
proximal blade margins entire;
juvenile blade (reddish or yellowish green), glabrous, puberulent, or sparsely pubescent abaxially, (hairs usually white, sometimes also ferruginous).
abaxial nectary absent, adaxial nectary oblong or narrowly oblong, 0.5–1.1 mm;
filaments distinct or connate less than 1/2 their lengths, glabrous, or hairy basally or on proximal 1/2;
anthers (yellow), ellipsoid, 0.4–0.6 mm.
abaxial nectary absent, adaxial nectary oblong, 0.2–1.4 mm, shorter than stipe;
stipe 2.1–3.2 mm;
ovary obclavate, often glaucous, glabrous, beak abruptly tapering to styles;
ovules 4–6 per ovary;
styles connate or distinct 1/2 their lengths, 0.1–0.2 mm;
stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.2–0.25–0.36 mm.
4–8 mm.
humboldtiana Willdenow: Humboldt willow is not known to occur in the flora area.
is characterized by: trees, 4–25 m;
branches highly brittle at base, bud-scale margins distinct and overlapping adaxially;
stipules on late leaves rudimentary or foliaceous;
largest medial leaf blade usually linear, abaxial surface not glaucous, adaxial dull; pistillate bract deciduous after flowering;
stamens 3–7;
capsules with distinct, often raised, white veins.;
it occurs throughout much of Mexico to central Chile.
staminate 11–21 × 4–8 mm, flowering branchlet 3–12 mm; pistillate loosely flowered, stout or subglobose, 14–37 × 5–14 mm, flowering branchlet 7–25 mm;
floral bract tawny or light rose, 0.8–1.6 mm, apex rounded, entire, abaxially very sparsely hairy distally, hairs straight or wavy.
= 38, 57, 76.
Salix humboldtiana
Salix pedicellaris
Salix humboldtiana is closely related to S. nigra in its generally narrow leaf blades, which are not glaucous abaxially. The two differ in the following characters: S. humboldtiana has leaf blades linear to sometimes narrowly oblong (10–28.6 times as long as wide), ovaries usually ovoid to ellipsoid, ovary walls often stomatiferous and with raised, white veins, and capsule valves relatively thick, slightly recurved. S. nigra has leaf blades usually narrowly lanceolate (6–13 times as long as wide), ovaries pyriform to obclavate, ovary walls neither stomatiferous nor notably veined, and capsule valves relatively thin and strongly recurved. Both species occur in Chihuahua, Mexico.
The report by R. I. Lonard et al. (1991) that specimens identified as Salix nigra from the lower Rio Grande, Texas, resemble S. humboldtiana in having strongly veined capsules suggests that S. humboldtiana, or intergrades with that species, may occur in Texas. Attempts to locate a voucher specimen were unsuccessful; because strongly veined capsules are diagnostic, further field study is indicated.
An earlier name, Salix chilensis Molina, has been applied to this species; it does not seem to pertain to this taxon (C. K. Schneider 1918).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Salix pedicellaris is found in Nunavut on Akimiski Island in James Bay and on the Belcher Islands in Hudson Bay.
Salix pedicellaris is very distinct with decumbent habit, leathery, glabrous leaves that are glaucous on both surfaces, loosely flowered catkins, ovaries reddish, glabrous and often glaucous, and stipes 2.1–3.2 mm. In the flora area, it hybridizes with six other species (see below). This compares with the closely related European S. myrtilloides Linnaeus, which is reported (B. Jonsell and T. Karlsson 2000+, vol. 1) to hybridize with seven species. The distinctive appearance may make hybrids easily recognizable but it is complex cytologically.
Hybrids:
Salix pedicellaris forms natural hybrids with S. arctica, S. argyrocarpa, S. athabascensis, S. chlorolepis, S. glauca var. cordifolia, S. pedicellaris, and S. pellita. Hybrids with S. candida and S. eriocephala have been reported (M. L. Fernald 1950) but no convincing specimens have been seen.
Salix pedicellaris × S. pellita (S. ×jamesensis Lepage) was described from the west coast of James Bay, Ontario. It is to be expected throughout northern Ontario and Quebec. According to E. Lepage (1964), its yellowish midveins and the hairiness of floral bracts superficially resemble those of S. pellita forma psila C. K. Schneider (syn. S. pellita forma denudata), but its reticulate leaf venation resembles that of S. pedicellaris.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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