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Humboldt's willow

Turnor's willow

Habit Shrubs, 1–2.5 m, (forming clones by layering).
Stems

branches yellow-brown or yellow-gray, not or weakly glaucous, (with sparkling wax crystals, dull or slightly glossy), pilose or villous;

branchlets gray-brown or red-brown, pubescent, villous, or velvety.

Leaves

stipules foliaceous, apex acute or convex;

petiole shallowly grooved adaxially, 4–13 mm, villous or pubescent adaxially;

largest medial blade (amphistomatous), narrowly oblong, narrowly elliptic, elliptic, oblanceolate, or lanceolate, 26–47 × 7.5–15 mm, 2.8–4.1 times as long as wide, base convex, rounded, or subcordate, margins slightly revolute or flat, serrate or serrulate, apex acuminate to acute, abaxial surface glaucous, glabrous, pilose, villous, or long-silky, hairs straight or wavy, adaxial dull, sparsely or moderately densely pilose or long-silky, especially on midrib;

proximal blade margins entire or serrulate;

juvenile blade reddish or yellowish green, sparsely to moderately densely long-silky or pubescent abaxially, hairs white.

Staminate flowers

adaxial nectary narrowly oblong or ovate, 0.7–1.1 mm;

filaments distinct or connate, hairy basally;

anthers yellow, 0.6–0.8 mm.

Pistillate flowers

adaxial nectary oblong, flask-shaped, or narrowly ovate, 0.4–1 mm, shorter than stipe;

stipe 2–4 mm;

ovary pyriform, glabrous, beaks slightly bulged below styles;

ovules 14–18 per ovary;

styles 0.3–0.5 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.16–0.23–0.28 mm.

Capsules

2.5–5 mm.

Salix

humboldtiana Willdenow: Humboldt willow is not known to occur in the flora area.

It

is characterized by: trees, 4–25 m;

branches highly brittle at base, bud-scale margins distinct and overlapping adaxially;

stipules on late leaves rudimentary or foliaceous;

largest medial leaf blade usually linear, abaxial surface not glaucous, adaxial dull;

pistillate bract deciduous after flowering;

stamens 3–7;

capsules with distinct, often raised, white veins.;

it occurs throughout much of Mexico to central Chile.

Catkins

flowering just before or as leaves emerge;

staminate stout, 16–30 × 8–10 mm, flowering branchlet 1.5–4 mm;

pistillate loosely flowered, stout, 18–22 × 9–11 mm, flowering branchlet 3–4 mm;

floral bract brown or tawny, 1.2–1.6 mm, apex acute or rounded, abaxially hairy throughout or proximally, hairs straight or wavy.

Salix humboldtiana

Salix turnorii

Phenology No flowering time data are available (probably May or Jun).
Habitat Active sand dunes
Elevation 200-300 m (700-1000 ft)
Distribution
from USDA
Mexico to central Chile
[BONAP county map]
from FNA
SK
[BONAP county map]
Discussion

Salix humboldtiana is closely related to S. nigra in its generally narrow leaf blades, which are not glaucous abaxially. The two differ in the following characters: S. humboldtiana has leaf blades linear to sometimes narrowly oblong (10–28.6 times as long as wide), ovaries usually ovoid to ellipsoid, ovary walls often stomatiferous and with raised, white veins, and capsule valves relatively thick, slightly recurved. S. nigra has leaf blades usually narrowly lanceolate (6–13 times as long as wide), ovaries pyriform to obclavate, ovary walls neither stomatiferous nor notably veined, and capsule valves relatively thin and strongly recurved. Both species occur in Chihuahua, Mexico.

The report by R. I. Lonard et al. (1991) that specimens identified as Salix nigra from the lower Rio Grande, Texas, resemble S. humboldtiana in having strongly veined capsules suggests that S. humboldtiana, or intergrades with that species, may occur in Texas. Attempts to locate a voucher specimen were unsuccessful; because strongly veined capsules are diagnostic, further field study is indicated.

An earlier name, Salix chilensis Molina, has been applied to this species; it does not seem to pertain to this taxon (C. K. Schneider 1918).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Salix turnorii is known from the Lake Athabasca sand dunes in northwestern Saskatchewan. Salix famelica in the Great Sand Hills, southern Saskatchewan, is very similar morphologically and may have been the source of populations ancestral to S. turnorii that moved into northern Saskatchewan during the warm Holocene Hypsithermal Period (ca. 9000–6000 yrs. B.P.).

Hybrids:

Salix turnorii forms natural hybrids with S. brachycarpa var. psammophila.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 34. FNA vol. 7, p. 122.
Parent taxa Salicaceae > Salix > subg. Protitea > sect. Humboldtianae Salicaceae > Salix > subg. Vetrix > sect. Cordatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. lutea var. turnorii
Name authority Willdenow Raup: J. Arnold Arbor. 17: 234, plate 193. (1936)
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