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halberd willow

rock willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Plants 0.2–1.5 m. Stems erect; branches brownish or red-brown, (dull), glabrous, long-silky, or villous to glabrescent; branchlets yellow-brown or gray-brown, long-silky, pilose, or moderately densely villous.
Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules absent or rudimentary;

petiole (shallowly to deeply grooved adaxially), 2–8 mm, (with 2 spherical glands distally, dark brown, sometimes basilaminar, sparsely pubescent or glabrous adaxially);

largest medial blade hypostomatous, (veins strongly impressed-reticulate), broadly elliptic, subcircular, or obovate, 18–67 × 10–40 mm, 1.1–2.3 times as long as wide, base rounded, convex, or subcordate, margins strongly revolute, crenate or subentire, apex rounded, convex, retuse, or toothed, abaxial surface sparsely to densely villous or long-silky, veins often with long, straight hairs, adaxial slightly glossy, glabrous or sparsely long-silky;

proximal blade margins entire or crenate;

juvenile blade (yellowish green), abaxially very densely long-silky.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

abaxial nectary 0.6–0.8 mm, adaxial nectary narrowly oblong, 0.5–1.2 mm, nectaries connate and shallowly cup-shaped, or distinct;

filaments distinct, hairy on proximal 1/2;

anthers ellipsoid or globose, 0.3–0.5 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

abaxial nectary (0–)0.8–0.9 mm, adaxial nectary oblong, ovate, or narrowly oblong to almost filiform, 0.7–1.4 mm, shorter to longer than stipe, nectaries distinct or connate and cup-shaped;

stipe 0.4–1.2 mm;

ovary pyriform or obnapiform, densely short-silky, hairs cylindrical, beak abruptly tapering to styles;

ovules 13–15 per ovary;

styles connate 1/2 their lengths to almost distinct, 0.2–0.4 mm;

stigmas flat, abaxially non-papillate with rounded tip, slenderly cylindrical, or 2 plump lobes, 0.2–0.28–0.36 mm.

Capsules

3.2–8 mm.

3–5 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

staminate 13–48 × 4–6.5(–8) mm, flowering branchlet 3–31(–50) mm; pistillate densely flowered, slender or stout, 18–56 × 4–10 mm, flowering branchlet 3–27(–40) mm;

floral bract tawny, 0.8–1.6 mm, apex rounded, entire, abaxially densely hairy, hairs straight.

2n

= 38.

= 38.

Salix hastata

Salix vestita

Phenology Flowering early Jun-late Jul. Flowering mid Jun-late Jul.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Moist to dry open forests and rocky streamsides, in upper montane and subalpine zones, rarely alpine
Elevation 0-1200 m (0-3900 ft) 0-2400 m (0-7900 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
MT; OR; WA; AB; BC; MB; NL; NS; NU; ON; QC; Asia (China [Xinjiang], Mongolia, Russia, e, c Siberia)
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix vestita is an ancient amphiberingian species. Its distribution includes a series of isolated, disjunct populations in Central Siberia, the northern Rocky Mountains, the west coast of Hudson Bay, and the northeastern arctic and subarctic. Occurrence in Nunavut is on Akpatok Island in Ungava Bay and on the Belcher Islands in Hudson Bay. It may be extirpated in Washington.

The flowering and vegetative branchlets sometimes have relatively short internodes. In subsequent years, branches have the appearance of short shoots similar to those in Alnus. Short shoots do not appear on all branches or in all years. The formation of short shoots may be related to adverse growing conditions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 64.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Chamaetia > sect. Chamaetia
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. leiolepis, S. vestita subsp. leiolepis, S. vestita var. psilophylla
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) Pursh: Fl. Amer. Sept. 2: 610. (1813)
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