The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

halberd willow

Tyrrell's willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Shrubs, 0.6–3.5 m, (sometimes forming clones by layering).
Stems

branches and branchlets red-brown, not to strongly glaucous, glabrous, (buds caprea-type or intermediate).

Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules rudimentary or foliaceous (often brownish) on early ones, foliaceous or rudimentary on late ones, (ca. 4 mm), apex acute;

petiole convex to flat, or shallowly grooved adaxially, 1–3.4–16 mm, glabrous, pilose, or short-silky adaxially;

largest medial blade (amphistomatous), narrowly elliptic, elliptic, oblanceolate, or obovate, 15–29–65 × 3.5–8.8–18 mm, 2.3–3.3–4.4 times as long as wide, base cuneate or convex, margins strongly to slightly revolute, entire, or very shallowly serrulate or shallowly serrulate-crenulate, apex acute, acuminate, or convex, abaxial surface glaucous, sparsely long-silky to glabrescent, hairs (ferruginous), straight, adaxial highly glossy, glabrous or sparsely short-silky (hairs white, sometimes also ferruginous);

proximal blade margins entire;

juvenile blade yellowish green or sometimes reddish, sparsely long-silky abaxially, hairs white and ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

adaxial nectary oblong, 0.8–1.1 mm;

filaments distinct, glabrous or hairy basally;

anthers purple turning yellow, ellipsoid, 0.4–0.7 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

adaxial nectary oblong or flask-shaped, 0.6–1.1 mm, equal to or shorter than stipe;

stipe 0.2–1 mm (–1.4 mm in cultivation);

ovary pyriform, long-silky, beak gradually tapering to styles;

ovules 12–16 per ovary;

styles 0.6–1.2 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.44–0.55–0.75 mm.

Capsules

3.2–8 mm.

3.6–5 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

flowering before leaves emerge; staminate stout, 14–35 × 12–16 mm, flowering branchlet 0 mm; pistillate densely flowered, stout, 17–51 × 10–13–22 mm, flowering branchlet 0–4 mm;

floral bract brown, black, or bicolor, 1–3.7 mm, apex acute to acuminate or rounded, abaxially hairy, hairs long, straight.

2n

= 38.

Salix hastata

Salix tyrrellii

Phenology Flowering early Jun-late Jul. Flowering mid Jun-mid Jul.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Active sand dunes, shrubby tundra
Elevation 0-1200 m (0-3900 ft) 200-600 m (700-2000 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
AB; NT; NU; SK
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix tyrrellii, first described from Lake Athabasca sand dunes in northwestern Saskatchewan and adjacent Alberta, is characterized by slender, amphistomatous leaves and relatively long, slender branchlets (G. W. Argus and J. W. Steele 1979). It probably evolved, in the past 10,000 years, from the widespread boreal S. planifolia. Originally, it was thought to be endemic to Lake Athabasca but recent collections from Nunavut and the Northwest Territories suggest that it may have a much wider range. Plants from Nunavut and the Northwest Territories do not have the long, slender branchlets found in sand dune populations, and identification is primarily based on the presence of amphistomatous leaves. This character, however, may be unreliable. For example, S. planifolia from 2610 m in the Glass Mountains, California, have amphistomatous leaves, suggesting that this character may be under environmental influence. The appropriate rank for this taxon remains uncertain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 140.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Vetrix > sect. Phylicifoliae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. planifolia subsp. tyrrellii
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) Raup: J. Arnold Arbor. 17: 231, plate 192. (1936)
Web links