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halberd willow

almond leaf willow, almond willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose.
Stems

branches glabrous or glabrescent;

branchlets yellow-brown, red-brown, or brownish, usually glabrous, rarely pilose.

Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules rudimentary to foliaceous on early ones (absent on proximal ones);

petiole deeply grooved adaxially, margins covering groove, 4–26 mm, pubescent or puberulent to glabrescent adaxially;

largest medial blade oblong, narrowly oblong, narrowly elliptic, elliptic, or lanceolate to obovate, 53–114 × 14–35 mm, 2.7–6.3 times as long as wide, base convex or cuneate, margins flat or slightly revolute, crenate or serrulate, apex acuminate, acute, or ± caudate, abaxial surface glabrous or glabrescent, adaxial dull or slightly glossy, glabrous or glabrescent;

proximal blade margins crenate or crenulate;

juvenile blade reddish.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

abaxial nectary 0.2–1.1 mm, adaxial nectary oblong, square, or ovate, 0.2–0.6 mm, distinct;

filaments distinct, hairy on proximal 1/2;

anthers ellipsoid.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

adaxial nectary obovate to square, 0.3–0.5 mm;

ovary pyriform, beak gradually tapering to or slightly bulged below styles;

ovules 30–36 per ovary;

styles distinct 1/2 their lengths, 0.2–0.3 mm;

stigmas flat, abaxially non-papillate with rounded tip, 0.1–0.2 mm.

Capsules

3.2–8 mm.

3–6 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

staminate 20–60 × 5.5–10 mm, flowering branchlet 3–17 mm; pistillate moderately to very densely flowered, slender to stout, 20–60 × 5–8 mm, flowering branchlet 5–31 mm;

floral bract 1–2.3 mm, apex rounded or acute, abaxially hairy (mainly proximally), hairs wavy.

2n

= 38.

= 38 (44), 57, or 88.

Salix hastata

Salix triandra

Phenology Flowering early Jun-late Jul. Flowering late spring.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Stream banks, waste places
Elevation 0-1200 m (0-3900 ft) 10-40 m (0-100 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
DC; ME; OH; VA; NS; ON; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix triandra usually has been overlooked in North American floras. At one time, it was a very important basket willow and probably was introduced into North America for that purpose. Some authors treat the glaucous and nonglaucous forms as subspecies (F. Martini and P. Paiero 1988; K. H. Rechinger 1993); A. K. Skvortsov (1999) noted that, although the two have somewhat distinct ranges, both kinds occur throughout the species and sometimes can be found in the same population. His suggestion that genetic inheritance of this character should be studied has not been taken up. The species is characterized by bark that is dark gray, smooth, and flaking in large irregular patches, as in Platanus ×acerifolia. The ovary-style transition is so indistinct that styles are often described as absent, but there are two, distinct styles, each terminating in a short stigma. A color change, later in the season, between the styles and ovary suggests that the tip of the ovary and the two distinct styles are both stylar tissues. In general, it appears that the styles are connate proximally and distinct distally.

Collections of Salix triandra made in 1934–35 by H. Hyland along the Penobscot River, Orono, Maine, were labeled by him as “introduced,” but they could have spread from cultivation or have been naturalized (A. Haines, pers. comm.). Recent naturalized occurrences are known from Toronto, Ontario, and Wolfville, Nova Scotia. Specimens identified as S. triandra by C. R. Ball are from Virginia and the District of Columbia. Salix triandra is reported to occur in Ohio (T. D. Sydnor and W. F. Cowen 2000) but voucher specimens were not found.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 50.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Salix > sect. Triandrae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. amygdalina, S. amygdalina var. discolor, S. triandra subsp. discolor
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) Linnaeus: Sp. Pl. 2: 1016. (1753)
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