Salix hastata |
Salix stolonifera |
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halberd willow |
creeping willow, sprouting leaf willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Plants 0.02–0.1 m, forming clones by layering or rhizomes. |
Stems | trailing or erect; branches red-brown, (sometimes weakly glaucous), glabrous; branchlets yellow-brown or greenish brown, glabrous. |
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Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules (sometimes marcescent) absent or rudimentary on early ones, rudimentary or foliaceous on late ones; petiole (deeply to shallowly grooved adaxially), 3–9–20 mm, (ciliate, glabrous adaxially); largest medial blade amphistomatous or hemiamphistomatous, (sometimes with 2 pairs of secondary veins arising at or close to base, arcing toward apex), elliptic, broadly elliptic, or subcircular, 16–42 × 12–30(–38) mm, 1–2 times as long as wide, base convex, cuneate, or rounded, margins flat or slightly revolute, entire or serrulate, ciliate (hairs wavy), apex convex, acuminate, rounded, or retuse, abaxial surface glabrous, adaxial highly glossy, glabrous; proximal blade margins entire or irregularly serrulate; juvenile blade pilose to glabrescent abaxially. |
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
abaxial nectary (0–)0.2–0.7 mm, adaxial nectary oblong, narrowly oblong, or ovate, 0.6–1.3 mm, nectaries distinct; filaments distinct (glabrous); anthers ellipsoid, 0.5–0.6 mm. |
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
abaxial nectary absent, adaxial nectary oblong, 0.5–1.4 mm, longer than stipe; stipe 0.2–0.8 mm; ovary pyriform, sometimes glaucous, glabrous, beak gradually tapering to styles; ovules 12–13 per ovary; styles connate to distinct 1/2 their lengths, (0.6–)0.8–2 mm; stigmas flat, abaxially non-papillate with rounded or pointed tip, or slenderly cylindrical, 0.32–0.5–0.88 mm. |
Capsules | 3.2–8 mm. |
4–10 mm. |
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
staminate 13–31 × 7–11 mm, flowering branchlet 1–15 mm; pistillate moderately densely to loosely flowered, stout to globose, 15–54(–90 in fruit) × 6–15 mm, flowering branchlet 2–42 mm; floral bract brown, 1.6–2 mm, apex rounded, entire, abaxially sparsely hairy throughout or distally, hairs straight or wavy. |
2n | = 38. |
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Salix hastata |
Salix stolonifera |
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Phenology | Flowering early Jun-late Jul. | Flowering early Jun-early Jul. |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | Arctic, subarctic, and alpine, wet sedge meadows, hummocky tundra, raised center polygons, Dryas-willow-sedge tundra, Dryas mats on dry ridge tops |
Elevation | 0-1200 m (0-3900 ft) | 0-1000 m (0-3300 ft) |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
AK; AB; BC |
Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Hybrids: Salix stolonifera forms natural hybrids with S. arctica and S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 75. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Chamaetia > sect. Ovalifoliae |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | Coville: Proc. Wash. Acad. Sci. 3: 333, plate 41, fig. 1. (1901) |
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