Salix hastata |
Salix sphenophylla |
|
---|---|---|
halberd willow |
wedge-leaf willow |
|
Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Plants 0.03–0.12 m, not clonal or forming clones by layering. |
Stems | trailing and rooting; branches yellow-brown or brownish, glabrous; branchlets yellow-brown, glabrous. |
|
Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules absent or rudimentary; petiole 4–25 mm, (glabrous or pilose adaxially); largest medial blade hypostomatous, narrowly elliptic, broadly elliptic, obovate, or very broadly obovate, 19–52 × 10–28 mm, 1–3 times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire, apex convex, retuse, or rounded, abaxial surface glabrous, pilose or sparsely long-silky to glabrescent, hairs straight or wavy, adaxial slightly glossy, glabrous or pilose; proximal blade margins entire; juvenile blade (reddish), very sparsely long-silky abaxially. |
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
abaxial nectary absent, adaxial nectary oblong, 0.6–1 mm; filaments distinct; anthers ellipsoid or shortly cylindrical, 0.4–0.6 mm. |
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
adaxial nectary oblong or ovate, 0.7–1.6 mm, equal to or longer than stipe; stipe 0.5–1.4 mm; ovary obclavate or pyriform, glabrous, or patchy or streaky pilose or villous, especially on beak, beak gradually tapering to or slightly bulged below styles; ovules 10–18 per ovary; styles 0.6–1.8 mm; stigmas flat, abaxially non-papillate with pointed tip, or slenderly cylindrical, 0.32–0.5–0.68 mm. |
Capsules | 3.2–8 mm. |
4–12 mm. |
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
staminate 21–53 × 7–13 mm, flowering branchlet 8–20 mm; pistillate loosely to densely flowered, slender or stout, 32–79 × 7–18 mm, flowering branchlet 4–27 mm; floral bract brown or black, 1.1–2 mm, apex rounded, entire, abaxially hairy or ciliate, hairs straight. |
2n | = 38. |
= 38, 57. |
Salix hastata |
Salix sphenophylla |
|
Phenology | Flowering early Jun-late Jul. | Flowering mid Jun-late Jul (early Aug). |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | Stony or gravelly substrates on talus, rocky outcrops, dry, stony tundra, sandy and moss tundra |
Elevation | 0-1200 m (0-3900 ft) | 10-900 m (0-3000 ft) |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
AK; NT; YT; e Asia (Chukotka, Russian Far East, e Siberia) |
Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The patchy or streaky indumentum on the ovaries of some plants suggests that they may be hybrids. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 82. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Chamaetia > sect. Diplodictyae |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | S. sphenophylla subsp. pseudotorulosa |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | A. K. Skvortsov: Spisok Rast. Gerb. Fl. S.S.S.R. Bot. Inst. Vsesoyuzn. Akad. Nauk 16: 62. (1966) |
Web links |