Salix hastata |
Salix raupii |
|
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halberd willow |
Raup willow, Raup's willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Plants 1.2–1.8 m, not clonal. |
Stems | erect; branches gray-brown, glabrous; branchlets yellow-brown, glabrous. |
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Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules foliaceous; petiole 5–9 mm; largest medial blade narrowly elliptic, 32–58 × 12–19 mm, 2–3.3 times as long as wide, base cuneate or convex, margins slightly revolute, entire, apex acute to acuminate, abaxial surface glabrous, adaxial slightly glossy, glabrous; proximal blade margins shallowly serrulate; juvenile blade glabrous. |
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
abaxial nectary 0.3–0.8 mm, adaxial nectary narrowly oblong, 0.6–1 mm, nectaries distinct; filaments distinct, glabrous; anthers ellipsoid, shortly cylindrical, or globose, 0.4–0.7 mm. |
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.5–1.1 mm, equal to or longer than stipe; stipe 0.4–1.2 mm; ovary pyriform, usually glabrous, rarely puberulent, beak slightly bulged below styles; ovules 12 per ovary; styles 0.6–0.8 mm; stigmas flat, abaxially not papillate with rounded tip, or broadly cylindrical, 0.3–0.5 mm. |
Capsules | 3.2–8 mm. |
4.4–8 mm. |
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
staminate 17.5–42 × 5–13 mm, flowering branchlet 6–7 mm; pistillate moderately densely flowered, stout, 20–40 × 6–12 mm, flowering branchlet 4–7 mm; floral bract tawny or bicolor, 1.3–2.5 mm, apex rounded, entire, abaxially glabrous. |
2n | = 38. |
|
Salix hastata |
Salix raupii |
|
Phenology | Flowering early Jun-late Jul. | Flowering late Jun. |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | Thickets in moist, open forests, gravel floodplains |
Elevation | 0-1200 m (0-3900 ft) | 800-1500 m (2600-4900 ft) |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
AB; BC; NT; YT |
Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Salix raupii resembles glabrous S. glauca var. villosa. Thin-layer chromatography of leaf phenolics in S. raupii revealed a pattern similar to those of S. glauca vars. villosa and acutifolia and S. athabascensis (G. W. Argus, unpubl.). Based on overall similarity, its nearest neighbors are S. glauca, in a broad sense, and S. athabascensis (Argus 1997). The sectional placement of this species is uncertain. It is placed here in sect. Myrtilloides because it clusters with S. athabascensis, but it is evidently close to S. glauca and may be a species of intersectional hybrid origin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 85. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Chamaetia > sect. Myrtilloides |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | Argus: Canad. J. Bot. 52: 1303, plate 1. (1974) |
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