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halberd willow

false mountain willow, serviceberry willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Plants 1–6 m. Stems: branches red-brown or yellow-brown, not or weakly glaucous, (slightly or highly glossy), glabrous or glabrescent; branchlets yellow-green, red-brown, or brownish, glabrous, pilose, or densely villous, (inner membranaceous bud-scale layer free, separating from outer layer).
Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules foliaceous, (prominent, 5–9–15 mm), apex rounded to acute, sometimes acuminate;

petiole (usually reddish), shallowly grooved, or convex to flat adaxially, 6–20 mm, short-silky or velvety adaxially;

largest medial blade broadly to narrowly elliptic, or ovate to broadly obovate, (25–)30–86(–118) × 12–51 mm, 1.4–3 times as long as wide, base convex, rounded, cuneate, subcordate, cordate, or sometimes irregularly lobed, margins flat, serrulate or crenate, apex acute, acuminate, or convex, abaxial surface glaucous, glabrous, pubescent, or pilose, hairs wavy, adaxial slightly glossy or dull, glabrous, puberulent, pubescent, or pilose, midrib hairy;

proximal blade margins entire or serrulate;

juvenile blade reddish, glabrous or pubescent abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

adaxial nectary oblong, 0.3–1 mm;

filaments distinct or connate less than 1/2 their lengths, glabrous;

anthers purple turning yellow, 0.4–0.5 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

adaxial nectary oblong, flask-shaped, 0.3–0.8 mm, shorter than stipe;

stipe 0.5–0.8–3 mm;

ovary pyriform or obclavate, glabrous, beak gradually tapering to styles;

ovules 18 per ovary;

styles (0.5–)0.7–1.8 mm;

stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.1–0.21–0.29 mm.

Capsules

3.2–8 mm.

4–7 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

flowering before leaves emerge; staminate stout, 16–39 × 10–12 mm, flowering branchlet 0 mm; pistillate densely or moderately densely flowered, slender to globose, 17–73 × 8–20 mm, flowering branchlet 0–5 mm;

floral bract brown or black, 1–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight.

2n

= 38.

= 38.

Salix hastata

Salix pseudomonticola

Phenology Flowering early Jun-late Jul. Flowering late Apr-early Jun.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Moist fens in drainageways in white spruce forests, treed bogs, balsam poplar forests, floodplains
Elevation 0-1200 m (0-3900 ft) 0-2500 m (0-8200 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
AK; ID; MN; MT; SD; WA; WY; AB; BC; MB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix pseudomonticola is characterized by precocious flowering; catkins sessile; juvenile leaf blades, petioles, and proximal midribs reddish; stipules prominent; and leaves and branchlets sparsely hairy. Branches older than two years have a distinctive pattern, which consists of a series of longitudinal splits in epidermis produced as the branch expands. The edge of epidermis around the split, where it has separated from the branch, is yellow and contrasts with the red-brown branch to which the epidermis still adheres.

Vegetative specimens of Salix pseudomonticola with yellow-brown branches can be confused with S. famelica. They may be separated by their juvenile leaf margins prominently and closely gland-dotted; stipules usually prominent, sometimes early deciduous; leaves broader (1.4–3 times as long as wide versus 2.6–7 in S. famelica); and petioles slender and often longer in relation to blade length. The possibility of hybridization needs study.

Vegetative specimens of Salix pseudomonticola can be distinguished from S. pyrifolia by juvenile leaves reddish and almost always with some ferruginous hairs, versus yellowish-green and glabrous or with white hairs, and mature leaves usually dull adaxially versus glossy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 117.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Vetrix > sect. Hastatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. barclayi var. pseudomonticola
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) C. R. Ball: Contr. U.S. Natl. Herb. 22: 321. (1921)
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