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halberd willow

polar willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Plants 0.01–0.09 m, (dwarf), forming clones by rhizomes.
Stems

erect;

branches red-brown, (often glaucous, dull), glabrous;

branchlets brownish, glabrous.

Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules absent or rudimentary;

petiole (deeply grooved), 1.3–10 mm, (glabrous adaxially);

largest medial blade (deciduous in autumn, 2 pairs of secondary veins arising at or close to base, arcing toward apex), elliptic, broadly elliptic, obovate, or subcircular, 5–32 × 7–18 mm, 1.1–2.8 times as long as wide, base convex, rounded, or cuneate, margins slightly revolute or flat, entire, ciliate, apex usually rounded or convex, sometimes retuse, abaxial surface (rarely glaucous), glabrous or pilose, adaxial slightly glossy, glabrous;

proximal blade margins entire;

juvenile blade glabrous.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

abaxial nectary 0.3–0.7 mm, adaxial nectary oblong, narrowly oblong, square, or ovate, 0.5–1.4 mm, nectaries distinct;

filaments usually distinct, sometimes connate proximally, glabrous;

anthers ellipsoid or ovoid, 0.4–0.6 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

abaxial nectaries absent, adaxial nectary narrowly oblong, oblong, or ovate, 0.8–1.8 mm, longer than stipe;

stipe 0.2–0.7 mm;

ovary obclavate or pyriform, densely villous to pilose, hairs flattened, beak gradually tapering to or slightly bulged below styles;

ovules 12–17 per ovary;

styles connate to distinct 1/2 their lengths, 0.7–1.2 mm;

stigmas flat, abaxially non-papillate with pointed tip, or slenderly to broadly cylindrical, 0.3–0.6(–0.7) mm.

Capsules

3.2–8 mm.

4.8–8.25 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

sometimes from subterminal buds; staminate 9–34 × 6–12 mm, flowering branchlet 1.5–14 mm; pistillate densely or moderately flowered (more than 15 flowers), stout to globose, 10–50 × 7–13 mm, flowering branchlet 1–12 mm;

floral bract brown, black, or bicolor, 1.5–2.5 mm, apex rounded or convex, entire, abaxially sparsely hairy, hairs straight or wavy, (exceeding bract by 0.6–1.12–1.8 mm).

2n

= 38.

= 76, 114.

Salix hastata

Salix polaris

Phenology Flowering early Jun-late Jul. Flowering mid Jun-early Aug.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Arctic-alpine, moist late snowbed and snowflush areas, talus and scree slopes, sides of depressed center frost polygons, sedge meadows, and mud boils, calcareous tills, sandy marine silts
Elevation 0-1200 m (0-3900 ft) 0-1800 m (0-5900 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
AK; BC; NT; NU; YT; Eurasia (Chukotka, Novaya Zemlya, Russian Far East, arctic Siberia, Spitzbergen, and Sweden)
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The sectional placement of Salix polaris is uncertain. It was placed in sect. Myrtosalix (G. W. Argus 1997) but more recently Argus et al. (1999) placed it in sect. Herbella. This polyploid species may be an intersectional hybrid.

Hybrids:

Salix polaris forms natural hybrids with S. arctica and perhaps S. rotundifolia.

Salix polaris × S. rotundifolia: This putative hybrid occurs in Alaska and the Yukon. Many plants previously identified as S. rotundifolia but that have ovaries with hairs on the beaks or in patches, leaves not commonly marcescent, and catkins with fewer flowers than in S. polaris may be this hybrid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 69.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Chamaetia > sect. Herbella
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. polaris subsp. pseudopolaris, S. polaris var. selwynensis
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) Wahlenberg: Fl. Lapp., 261, plate 13, fig. 1. (1812)
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