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halberd willow

meadow willow, skeleton-leaf willow, slender willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Plants 1–6 m. Stems: branches red-brown or violet, not or weakly glaucous, (dull or slightly glossy), puberulent; branchlets yellow-green to red-brown, sparsely pubescent or moderately densely velvety, (buds alba-type or intermediate).
Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules rudimentary or absent;

petiole shallowly grooved adaxially, 3–11 mm, pubescent, or velvety to glabrescent adaxially;

largest medial blade lorate or very narrowly elliptic, 38–110 × 6–19 mm, 5–9 times as long as wide, base cuneate or convex, margins flat to slightly revolute, entire, serrate, serrulate, or spinulose-serrate, apex acute to acuminate, abaxial surface glaucous, densely long-silky to glabrescent, hairs (white, sometimes also ferruginous), adaxial dull or slightly glossy, glabrous or sparsely pubescent, (hairs white, sometimes also ferruginous);

proximal blade margins sometimes serrulate;

juvenile blade moderately densely long-silky abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

adaxial nectary square, ovate, or oblong, 0.3–0.7 mm;

filaments distinct, hairy basally;

anthers purple turning yellow, ellipsoid or globose, 0.4–0.6 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

adaxial nectary oblong to ovate, 0.3–0.9 mm;

stipe 1.5–4 mm;

ovary pyriform, beak abruptly tapering to styles;

ovules 6–12 per ovary;

styles 0–0.5 mm;

stigmas slenderly to broadly cylindrical, 0.26–0.4–0.8 mm.

Capsules

3.2–8 mm.

5–9 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

flowering as leaves emerge; staminate stout to globose, 12–29 × 6–17 mm, flowering branchlet 0.8–3 mm; pistillate loosely flowered, stout to globose, 12–39 × 6–18 mm, flowering branchlet 1–11 mm;

floral bract brown, tawny, light rose, or bicolor, 1–2 mm, apex rounded, abaxially sparsely hairy, hairs straight.

2n

= 38.

= 38.

Salix hastata

Salix petiolaris

Phenology Flowering early Jun-late Jul. Flowering mid Apr-mid Jun.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Sedge meadows, openings in moist, low, rich deciduous woods, sandy or peaty wet prairies, lakeshores
Elevation 0-1200 m (0-3900 ft) 10-2700 m (0-8900 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
CO; CT; IA; IL; IN; MA; ME; MI; MN; ND; NE; NH; NJ; NY; OH; PA; SD; VT; WI; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

See 107. Salix sericea for a comparative description.

Because reproductive barriers between Salix petiolaris and S. eriocephala are weak, A. Mosseler (1990) suggested that their morphological variability may be due to interspecific gene flow.

Hybrids:

Salix petiolaris forms natural hybrids with S. bebbiana, S. candida, S. eriocephala, S. famelica, S. pellita, and S. sericea. Hybrids with S. alba have been reported (M. L. Fernald 1950) but no convincing specimens have been seen. Controlled pollinations with S. discolor produced no seed (A. Mosseler 1990).

Reports of Salix petiolaris × S. sericea from Massachusetts and Pennsylvania (C. K. Schneider 1921) probably refer to the densely sericeous variant of S. petiolaris. It is sometimes named S. ×subsericea (Andersson) C. K. Schneider but does not seem to be a hybrid (G. W. Argus 1965, 1986; E. G. Voss 1972–1996, vol. 2).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 152.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Vetrix > sect. Geyerianae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. gracilis, S. gracilis var. textoris, S. petiolaris var. gracilis, S. ×subsericea
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) Smith: Trans. Linn. Soc. 6: 122. (1802)
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