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halberd willow

satiny willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Shrubs, 0.5–6 m, (sometimes forming clones by stem fragmentation).
Stems

branches (highly to ± brittle at base), red-brown, violet or yellow-brown, usually strongly glaucous, glabrescent;

branchlets yellow-brown or red-brown, (usually strongly glaucous), glabrous or densely to sparsely pubescent or tomentose, (buds caprea-type).

Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules absent or rudimentary on early ones, rudimentary, absent, or foliaceous on late ones;

petiole convex to flat, or shallowly grooved adaxially, 3–6.3–14 mm, (sometimes dark spherical glands distally), glabrous or pubescent adaxially;

largest medial blade linear, lorate, or narrowly elliptic 40–79–123 × 6–12–20 mm, (2.3–)4.2–7.2–11.3 times as long as wide, base convex or cuneate, margins strongly or slightly revolute, entire, sinuate or sometimes crenulate, (glands submarginal or epilaminal), apex acuminate to acute, abaxial surface glaucous (sometimes obscured by hairs), densely villous, short-silky, woolly, or tomentose to glabrescent, hairs (white, sometimes also ferruginous), straight or wavy, adaxial slightly to highly glossy, glabrous, sparsely villous or pubescent (hairs white, sometimes also ferruginous);

proximal blade margins entire;

juvenile blade reddish or yellowish green, densely tomentose, short-silky, pubescent, or glabrous abaxially, hairs white, sometimes also ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

adaxial nectary oblong or narrowly oblong, 0.6–1 mm;

filaments distinct, glabrous or hairy basally;

anthers purple turning yellow, ellipsoid, 0.4–0.6 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

adaxial nectary oblong to depressed-ovate, 0.3–1 mm, shorter than or equal to stipe;

stipe 0.5–1.1 mm;

ovary pyriform, short-silky, beak sometimes slightly bulged below styles;

ovules 10–18 per ovary;

styles 0.6–1.5 mm;

stigmas slenderly cylindrical, 0.4–0.55–0.76 mm.

Capsules

3.2–8 mm.

3.5–6.5 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

flowering before leaves emerge; staminate stout, 20–39 × 7–20 mm, flowering branchlet 0–2 mm; pistillate densely flowered, slender, stout, or subglobose, 19–65(–80 in fruit) × 7–17 mm, flowering branchlet 0–7 mm;

floral bract tawny, brown, or black, 1–2.6 mm, apex acute, convex, or rounded, abaxially hairy, hairs straight.

2n

= 38.

= 38.

Salix hastata

Salix pellita

Phenology Flowering early Jun-late Jul. Flowering late Apr-late Jun.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Sandy or gravelly floodplains, stream and lake margins, marshes, fens, coastal dunes, metamorphic or calcareous substrates
Elevation 0-1200 m (0-3900 ft) 0-800 m (0-2600 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
ME; MI; MN; NH; VT; WI; MB; NB; NL; NS; ON; QC; SK; SPM
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix pellita sometimes has foliaceous stipules on late leaves. When present, they are correlated with hairy rather than glabrous branchlets. There is no clear evidence of hybridity in such specimens but further study is indicated.

Vegetative specimens of Salix pellita can be difficult to separate from S. viminalis and S. ×smithiana in eastern Canada, where the latter were introduced for coarse basketry and have become naturalized. The introduced species usually are tall shrubs to small trees, with branches usually flexible at base and not strongly glaucous, and their leaves tend to be broader. In contrast, S. pellita is a mid shrub rarely to 6 m but never tree-like, its branches usually are highly brittle at the base and often strongly glaucous, and its leaves tend to be narrower.

See 85. Salix drummondiana for further comparative descriptions.

Hybrids:

Salix pellita forms natural hybrids with S. alaxensis var. alaxensis, S. discolor, S. pedicellaris, S. petiolaris, and S. planifolia.

Salix pellita × S. petiolaris: Leaves of this hybrid are distinctly serrate and flat, as in S. petiolaris, but branchlets are glaucous and bud gradation is caprea-type as in S. pellita. It is uncommon in eastern Saskatchewan.

Salix pellita × S. planifolia: This cross is suspected to occur in Labrador, Newfoundland, Quebec, and Ontario. Both parents are tetraploids and flower at the same time. The name S. pellita forma psila may apply to this hybrid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 136.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Vetrix > sect. Phylicifoliae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. chlorophylla
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) (Andersson) Bebb: Bot. Gaz. 16: 106. (1891)
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