Salix hastata |
Salix ovalifolia |
|||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
halberd willow |
arctic seashore or oval-leaf willow, oval-leaf willow |
|||||||||||||
Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Plants 0.02–0.05 m, not clonal or forming clones by layering. | ||||||||||||
Stems | trailing; branches yellow-brown, gray-brown, or red-brown, glabrous or hairy; branchlets yellow-green, yellow-brown, or red-brown, glabrous or pilose. |
|||||||||||||
Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules usually absent or rudimentary, rarely foliaceous; petiole (deeply to shallowly grooved adaxially), 1.1–16 mm, (glabrous); largest medial blade hypostomatous, narrowly to broadly elliptic, circular, subcircular, or obovate, 13–46 × 7–20 mm, 1–3.4 times as long as wide, base subcordate, cordate, rounded, or convex, margins slightly revolute or flat, entire, sometimes ciliate, apex convex, rounded, acuminate, acute, or retuse, abaxial surface glabrous, villous, long-silky, pubescent, or pilose, hairs wavy or straight, adaxial highly glossy, usually glabrous; proximal blade margins entire; juvenile blade (reddish or yellowish green), pilose, villous, or long-silky abaxially. |
||||||||||||
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
abaxial nectary 0.6–1 mm, adaxial nectary oblong or ovate, 0.6–1.6 mm, nectaries distinct or connate and cup-shaped; filaments distinct or connate less than 1/2 their lengths (glabrous); anthers elliptic, short-cylindrical, or globose, 0.3–0.5(–0.6) mm. |
||||||||||||
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
abaxial nectary (0–)0.4–0.6 mm, adaxial nectary longer than stipe; stipe 0.2–1.4 mm; ovary obclavate or pyriform, glaucous or not, usually glabrous or tomentose, sometimes pubescent or villous, beak abruptly tapering to styles; ovules 10–15 per ovary; styles 0.2–0.8 mm; stigmas flat, abaxially non-papillate with pointed tip, or slenderly cylindrical, 0.32–0.41–0.64 mm. |
||||||||||||
Capsules | 3.2–8 mm. |
5.2–9.6 mm. |
||||||||||||
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
staminate 4.8–46 × 5–11 mm, flowering branchlet 1.5–24 mm; pistillate moderately densely flowered, stout, subglobose, globose, or slender, 6.3–50 × 5–28 mm, flowering branchlet 2.5–22 mm; floral bract brown, greenish, or bicolor, 1.2–2 mm, apex rounded, entire or, sometimes, 2-fid, abaxially hairy, hairs straight or wavy. |
||||||||||||
2n | = 38. |
= 38. |
||||||||||||
Salix hastata |
Salix ovalifolia |
|||||||||||||
Phenology | Flowering early Jun-late Jul. | |||||||||||||
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | |||||||||||||
Elevation | 0-1200 m (0-3900 ft) | |||||||||||||
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
AK; NT; YT; Asia |
||||||||||||
Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). The varieties of Salix ovalifolia are relatively minor variants; their ranges overlap and their differences in leaf shape and ovary indumentum intergrade (G. W. Argus 1969, 1973). The only one with a more or less distinctive geographical distribution is var. cyclophylla; where its range overlaps with var. ovalifolia there is intergradation. Variety arctolitoralis, which is characterized by larger leaves and catkins, may be an ecotype. Variety glacialis is known only from near Point Barrow, Alaska. E. Hultén (1968) suggested, probably based on its often tomentose ovaries, that it is S. arctica × S. ovalifolia. All varieties of the species have some plants with hairy ovaries, but the suggestion that this character is an indication of hybridization deserves study. Hybrids: Salix ovalifolia forms natural hybrids with S. arctica and S. fuscescens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||
Key |
|
|||||||||||||
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 77. | ||||||||||||
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Chamaetia > sect. Ovalifoliae | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | |||||||||||||
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | Trautvetter: Nouv. Mém. Soc. Imp. Naturalistes Moscou 2: 306, plate 13. (1832) | ||||||||||||
Web links |