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halberd willow

bayberry willow, blue-leaf willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Plants 0.3–5 m, (sometimes forming clones by stem fragmentation or layering).
Stems

branches (sometimes highly brittle at base), red-brown or yellow-brown, not or weakly glaucous, (slightly or highly glossy), glabrous or villous;

branchlets red-brown or yellow-brown, glabrous or sparsely to very densely villous, (buds caprea-type, inner membranaceous bud-scale layer free and separating or not).

Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules rudimentary or foliaceous on early ones, foliaceous on late ones, apex acute or acuminate;

petiole shallowly grooved, or convex to flat adaxially, 3.5–7.3–13 mm, (sometimes with 2 spherical glands distally), villous, tomentose, pilose, or pubescent adaxially;

largest medial blade narrowly oblong, narrowly elliptic, elliptic, or oblanceolate, 35–61.3–110 × 11–16–46 mm, 2–2.7–5.2 times as long as wide, base convex, rounded, subcordate, or cuneate, margins flat or slightly revolute, (thickened and raised), crenulate or serrulate, apex acuminate, acute, or convex, abaxial surface usually very thickly glaucous, glabrous or pilose, midribs pubescent to tomentose, hairs (white, often also ferruginous), curved, wavy, or straight, adaxial slightly glossy, glabrous or pilose, midribs sparsely pubescent (hairs white, sometimes also ferruginous);

proximal blade margins entire or serrulate;

juvenile blade translucent, reddish or yellowish green, glabrous or sparsely pubescent abaxially, midribs often densely hairy, hairs white, sometimes also ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

adaxial nectary narrowly oblong, oblong, square, or ovate, 0.44–0.48–1.4 mm;

filaments distinct, glabrous;

anthers yellow, (ellipsoid or globose), 0.52–0.69–0.76 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

adaxial nectary oblong, narrowly oblong, square, or flask-shaped, 0.56–0.8–1.4 mm, shorter than stipe;

stipe 0.96–1.7–3.4 mm;

ovary pyriform, glabrous, beak slightly bulged below styles;

ovules 12–14 per ovary;

styles (sometimes distinct), 0.3–0.8–1.3 mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, or slenderly cylindrical, 0.24–0.43–0.56 mm.

Capsules

3.2–8 mm.

5–7–11 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

staminate flowering before leaves emerge, pistillate as leaves emerge; staminate stout or slender, 23.5–35.6–51 × 9–12.7–22 mm, flowering branchlet 1–3.9–10 mm; pistillate loosely flowered, stout or slender, 19–42–62(–85 in fruit) × 8–13–18 mm, flowering branchlet 1.5–5.9–13 mm;

floral bract brown or bicolor, 1.2–1.8–3 mm, apex rounded or acute, sometimes toothed, abaxially hairy, hairs straight or wavy.

2n

= 38.

Salix hastata

Salix myricoides

Phenology Flowering early Jun-late Jul. Flowering early Apr-early Jul.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Stream and lake shores, gravel bars, subalpine conifer forests, alkaline fens, sea cliffs, dry limestone talus, swamps, tidal meadows, sand dunes
Elevation 0-1200 m (0-3900 ft) 0-1100 m (0-3600 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
IL; IN; ME; MI; OH; PA; WI; NB; NF; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants with densely villous branchlets and branches have been named var. albovestita. Branchlet indumentum varies widely in the species and seems to be continuous, with both villous and glabrous variants sometimes occurring in the same area. This characteristic is more common in populations on the shores of the Great Lakes and on the western coast of James Bay, but even these populations are variable.

Reports of Salix myricoides from Akimiski Island, Nunavut, were based on misidentified S. planifolia. See 68. S. eriocephala for differences.

Hybrids:

Salix myricoides forms natural hybrids with S. bebbiana, S. discolor, and S. glauca var. cordifolia. Hybrids with S. eriocephala have been reported (M. L. Fernald 1950) but no convincing specimens have been seen. Inasmuch as S. eriocephala and S. myricoides are very similar, hybrids between the two would be very difficult to identify.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 119.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Vetrix > sect. Hastatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei S. glaucophylla var. albovestita, S. glaucophylloides, S. glaucophylloides var. albovestita, S. glaucophylloides var. glaucophylla, S. myricoides var. albovestita
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) Muhlenberg: Ges. Naturf. Freunde Berlin Neue Schriften 4: 235, plate 6, fig. 2. (1803)
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