Salix hastata |
Salix ligulifolia |
|
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halberd willow |
strap-leaf willow, strap-leafed willow, tongue-leaf willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Shrubs, 1–8 m. Stems: branches yellow-brown, gray-brown, or red-brown, not glaucous, glabrous or villous; branchlets yellow-green or yellow-brown, glabrous, sparsely to densely villous, or velvety, (inner membranaceous bud-scale layer free, not separating from outer layer). |
Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules foliaceous, apex rounded, convex, acute or acuminate; petiole convex to flat, or shallowly grooved adaxially, 3–18 mm, glabrous, pilose, or velvety to glabrescent adaxially; largest medial blade lorate, narrowly oblong, or narrowly elliptic, 60–133 × 12–30 mm, 2.9–6.4 times as long as wide, base rounded, convex, or subcordate, margins flat, usually serrulate or serrate, rarely (apparently) entire, apex acuminate to acute, abaxial surface glaucous, glabrous, sparsely short-silky or pubescent, hairs straight or wavy, adaxial dull, glabrous, sparsely short-silky, or midrib pubescent; proximal blade margins entire, serrulate, or crenulate; juvenile blade reddish or yellowish green, glabrous or sparsely to moderately densely pilose or puberulent abaxially, hairs white. |
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
adaxial nectary narrowly oblong, oblong, flask-shaped, or triangular, 0.3–0.8 mm; filaments distinct or connate less than 1/2 their lengths (or appearing as a single stamen), glabrous or hairy basally; anthers purple or red turning yellow, (ellipsoid or globose), 0.5–0.8 mm. |
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
adaxial nectary oblong, narrowly oblong, or flask-shaped, 0.3–0.9 mm, shorter than stipe; stipe 0.9–2.5 mm; ovary pyriform, glabrous, beak sometimes slightly bulged below styles; ovules 12–21 per ovary; styles 0.2–0.6 mm; stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.16–0.25–0.4 mm. |
Capsules | 3.2–8 mm. |
4–6 mm. |
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
flowering as or just before leaves emerge; staminate stout, 20.5–34 × 8–11 mm, flowering branchlet 0–3 mm; pistillate moderately densely flowered, slender to subglobose, 15.5–49 × 8–18 mm, flowering branchlet 1–6 mm; floral bract brown or bicolor, 0.8–1.6 mm, apex acute or rounded, abaxially hairy throughout or proximally (hairs usually arising from rachis), hairs wavy or curly. |
2n | = 38. |
= 38. |
Salix hastata |
Salix ligulifolia |
|
Phenology | Flowering early Jun-late Jul. | Flowering late Mar-mid Jun(-late Jul). |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | Banks and floodplains, cienegas, sandy-clay or gravelly substrates |
Elevation | 0-1200 m (0-3900 ft) | 0-3100 m (0-10200 ft) |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
AZ; CA; CO; NM; OR; UT; WY
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Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Salix ligulifolia sometimes has leaf teeth that are so short that it is referred to as “entire-leaved.” That condition is uncommon and, even when some leaves appear to be entire, others with fine serrulations can be found on the same plant. Hybrids: Salix ligulifolia forms natural hybrids with S. geyeriana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 122. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Vetrix > sect. Cordatae |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | S. lutea var. ligulifolia, S. eriocephala var. ligulifolia |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | (C. R. Ball) C. R. Ball ex C. K. Schneider: J. Arnold Arbor. 2: 188. (1922) |
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