Salix hastata |
Salix jepsonii |
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halberd willow |
Jepson's willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Shrubs, 1–3 m, (sometimes forming clones by stem fragmentation). |
Stems | branches (highly brittle at base), yellow-brown or red-brown, weakly glaucous or not, (dull or slightly glossy), velvety or short-silky to glabrescent; branchlets gray-brown or red-brown, sparsely or moderately densely short-silky or velvety, (buds caprea-type or intermediate). |
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Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules usually rudimentary, sometimes foliaceous on early ones, foliaceous on late ones, apex acute; petiole shallowly grooved, or convex to flat adaxially, 3–12 mm, short-silky or velvety adaxially; largest medial blade lorate, narrowly oblanceolate, or oblanceolate, 43–74–103 × 8–25 mm, 2.9–4.8–7.7 times as long as wide, base convex or cuneate, margins slightly revolute to flat, entire, apex acuminate, convex, or acute, abaxial surface glaucous (sometimes obscured by hairs), densely short-silky, hairs straight, adaxial dull (sometimes thinly glaucous) to slightly glossy, sparsely short- or long-silky; proximal blade margins entire; juvenile blade yellowish green or reddish, densely long-silky abaxially, hairs white. |
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
adaxial nectary narrowly oblong to oblong, 0.4–0.8 mm; stamens 1 or 2; filaments distinct or connate less than 1/2 their lengths, hairy on proximal 1/2; anthers purple turning yellow, ellipsoid or shortly cylindrical, 0.6–0.8 mm. |
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
adaxial nectary oblong to narrowly oblong or flask-shaped, 0.3–0.6 mm, shorter than or equal to stipe; stipe 0.4–1.2 mm; ovary pyriform, densely long-silky, beak gradually tapering to styles; ovules 13–18 per ovary; styles 0.4–0.6 mm; stigmas flat, abaxially non-papillate with rounded to pointed tip, 0.2–0.3 mm. |
Capsules | 3.2–8 mm. |
3–5 mm. |
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
flowering as leaves emerge; staminate stout or subglobose, 16–16.5 × 11–13 mm, flowering branchlet 2.5–3 mm; pistillate moderately densely flowered, slender to stout, 33–55 × 10–11 mm, flowering branchlet 1.5–7 mm; floral bract brown or tawny, 0.8–2 mm, apex rounded or acute, abaxially hairy, hairs straight. |
2n | = 38. |
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Salix hastata |
Salix jepsonii |
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Phenology | Flowering early Jun-late Jul. | Flowering mid-late Jun. |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | Margins of lakes and streams, wet meadows, gravel, rocky, or bouldery substrates, granite |
Elevation | 0-1200 m (0-3900 ft) | 1000-3400 m (3300-11200 ft) |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
CA; NV; OR
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Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. D. Dorn (2000) justified treating Salix jepsonii as S. sitchensis var. angustifolia based on specimens of S. jepsonii from southern Oregon. Salix jepsonii does occur in southern Oregon but is distinct enough from S. sitchensis to be treated as a species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 160. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Vetrix > sect. Sitchenses |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | S. sitchensis var. angustifolia, S. pellita var. angustifolia, S. sitchensis var. ralphiana |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | C. K. Schneider: J. Arnold Arbor. 1: 89. (1919) |
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