Salix hastata |
Salix humboldtiana |
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halberd willow |
Humboldt's willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | |
Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
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Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
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Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
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Capsules | 3.2–8 mm. |
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Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
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Salix | humboldtiana Willdenow: Humboldt willow is not known to occur in the flora area. |
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It | is characterized by: trees, 4–25 m; branches highly brittle at base, bud-scale margins distinct and overlapping adaxially; stipules on late leaves rudimentary or foliaceous; largest medial leaf blade usually linear, abaxial surface not glaucous, adaxial dull; pistillate bract deciduous after flowering; stamens 3–7; capsules with distinct, often raised, white veins.; it occurs throughout much of Mexico to central Chile. |
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2n | = 38. |
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Salix hastata |
Salix humboldtiana |
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Phenology | Flowering early Jun-late Jul. | |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | |
Elevation | 0-1200 m (0-3900 ft) | |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
Mexico to central Chile |
Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Salix humboldtiana is closely related to S. nigra in its generally narrow leaf blades, which are not glaucous abaxially. The two differ in the following characters: S. humboldtiana has leaf blades linear to sometimes narrowly oblong (10–28.6 times as long as wide), ovaries usually ovoid to ellipsoid, ovary walls often stomatiferous and with raised, white veins, and capsule valves relatively thick, slightly recurved. S. nigra has leaf blades usually narrowly lanceolate (6–13 times as long as wide), ovaries pyriform to obclavate, ovary walls neither stomatiferous nor notably veined, and capsule valves relatively thin and strongly recurved. Both species occur in Chihuahua, Mexico. The report by R. I. Lonard et al. (1991) that specimens identified as Salix nigra from the lower Rio Grande, Texas, resemble S. humboldtiana in having strongly veined capsules suggests that S. humboldtiana, or intergrades with that species, may occur in Texas. Attempts to locate a voucher specimen were unsuccessful; because strongly veined capsules are diagnostic, further field study is indicated. An earlier name, Salix chilensis Molina, has been applied to this species; it does not seem to pertain to this taxon (C. K. Schneider 1918). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 34. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Protitea > sect. Humboldtianae |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | Willdenow |
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