Salix hastata |
Salix exigua |
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halberd willow |
coyote willow, narrow-leaf willow, sandbar willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Shrubs or trees, 0.5–5(–17) m. Stems: branches gray-brown, red-brown, or yellow-brown, villous, or tomentose to glabrescent; branchlets yellowish, yellow-brown, or red-brown, pubescent or puberulent, tomentose, short- or long-silky villous. | ||||
Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules absent or rudimentary on early ones, foliaceous or rudimentary on late ones; petiole 1–5(–10) mm, pubescent or puberulent, villous, or short-silky adaxially; largest medial blade (sometimes hypostomatous), linear or lorate, (glands marginal), 30–136–143 × 2–14 mm, 6.5–28(–37.5) times as long as wide, base cuneate, margins slightly revolute, entire or remotely spinulose-serrulate, apex acuminate or acute, abaxial surface glaucous (sometimes obscured by hairs), densely long-silky, villous or pilose to glabrescent, hairs appressed or spreading, straight or wavy, adaxial slightly glossy, sparsely or densely long-silky to glabrescent; proximal blade margins entire; juvenile blade yellowish green, densely long-silky-villous abaxially. |
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Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
abaxial and adaxial nectaries present and distinct; filaments hairy on proximal 1/2; anthers (sometimes reddish turning yellow). |
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Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
adaxial nectary oblong, ovate, or flask-shaped, relative adaxial nectary/stipe length variable even within same catkin; stipe 0–0.9 mm; ovary obclavate or pyriform, glabrous, pilose, or villous, or sometimes beak hairy, beak bulged below or abruptly tapering to styles; ovules 12–30 per ovary; styles (sometimes distinct), 0–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.1–0.5 mm. |
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Capsules | 3.2–8 mm. |
4–8 mm. |
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Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
staminate 7–54 × 2–10 mm, flowering branchlet 1.5–56 mm; pistillate loosely to densely flowered, slender or stout, 14.5–70 × 3–12 mm, flowering branchlet 2–55 mm; floral bract 1.2–2.6 mm, apex acute, convex, or rounded, entire or erose, hairs wavy, straight, or crinkled, abaxially hairy throughout or proximally, or glabrate. |
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2n | = 38. |
= 38. |
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Salix hastata |
Salix exigua |
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Phenology | Flowering early Jun-late Jul. | |||||
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | |||||
Elevation | 0-1200 m (0-3900 ft) | |||||
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
AZ; CA; CO; ID; MT; NE; NM; NV; OK; OR; TX; UT; WA; WY; AB; BC
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Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 54. | ||||
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Longifoliae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | S. fluviatilis var. exigua, S. longifolia var. exigua | ||||
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | Nuttall: N. Amer. Sylv. 1: 75. (1842) | ||||
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