Salix hastata |
Salix delnortensis |
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halberd willow |
Del Norte willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Shrubs, 1–2 m, (sometimes forming clones by stem fragmentation). |
Stems | branches (highly brittle at base), red-brown, not glaucous, tomentose or velvety to glabrescent; branchlets red-brown or yellow-brown, densely velvety, (buds caprea-type). |
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Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute; petiole convex to flat, or shallowly grooved adaxially, 6–16 mm, velvety or tomentose adaxially; largest medial blade elliptic or obovate, 53–102 × 29–54 mm, 1.3–2.8 times as long as wide, base cuneate or convex, margins slightly revolute, entire or sinuate, apex convex, rounded, or acute, abaxial surface glaucous (sometimes obscured by hairs), densely to sparsely velvety, tomentose, villous, or short-silky, hairs erect or spreading, wavy, adaxial dull, sparsely tomentose or short-silky; proximal blade margins entire; juvenile blade green, very densely velvety or long-silky abaxially, hairs white or gray. |
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
adaxial nectary narrowly oblong to oblong, 0.5–1 mm; filaments distinct or slightly connate, glabrous; anthers purple turning yellow, ellipsoid, 0.6–0.7 mm. |
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
adaxial nectary narrowly oblong to oblong, 0.4–0.9 mm, longer than stipe; stipe 0–0.3 mm; ovary obturbinate, short-silky or densely pubescent, beak gradually tapering to styles; ovules 14–18 per ovary; styles 0.6–1.2 mm; stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.2–0.32–0.4 mm. |
Capsules | 3.2–8 mm. |
4 mm. |
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
flowering before leaves emerge; staminate stout, 25–30 × 6–13 mm, flowering branchlet 0–5 mm; pistillate moderately densely flowered, slender to stout, 17–53 × 6–8 mm, flowering branchlet 1–3 mm; floral bract brown, 1.2–2.2 mm, apex rounded, abaxially hairy, hairs straight. |
2n | = 38. |
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Salix hastata |
Salix delnortensis |
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Phenology | Flowering early Jun-late Jul. | Flowering late Mar-early May. |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | Streamshores, gravel to boulder substrates, serpentine soils |
Elevation | 0-1200 m (0-3900 ft) | 90-500 m (300-1600 ft) |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
CA; OR
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Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The origin of the serpentine endemics Salix breweri and S. delnortensis is still unresolved (G. W. Argus 1997; R. D. Dorn 2000). The possibility that they are of hybrid origin, perhaps involving S. lasiolepis and S. sitchensis (Dorn 1976), cannot be resolved by field and herbarium studies alone; cytological and experimental methods need to be employed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 161. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Vetrix > sect. Sitchenses |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | S. breweri var. delnortensis |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | C. K. Schneider: J. Arnold Arbor. 1: 96. (1919) |
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