Salix hastata |
Salix atrocinerea |
|
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halberd willow |
large gray willow, rusty willow |
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Habit | Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. | Shrubs, 3–12 m. Stems: branches yellow-brown, gray-brown, or red-brown, not glaucous, pilose or villous to glabrescent, (peeled wood often with many striae, to 45 mm); branchlets gray-brown or yellow-brown, puberulent, pilose, villous, or velvety. |
Leaves | stipules foliaceous, apex acute to acuminate; petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially; largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous); proximal blade margins entire or finely serrulate; juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous. |
stipules (sometimes marcescent) foliaceous, apex acute; petiole convex to flat adaxially, 3–15 mm, tomentose, or velvety to glabrescent adaxially; largest medial blade narrowly to broadly elliptic, oblanceolate, obovate, broadly obovate, 29–105 × 14–52 mm, 1.8–4.3 times as long as wide, base cuneate or convex, margins slightly revolute, entire, crenate, or sinuate, (glands submarginal or epilaminal), apex acute, convex, or acuminate, abaxial surface glaucous, tomentose or coarsely villous to glabrescent, hairs (white, sometimes also ferruginous), erect, spreading, or appressed, wavy or curved, adaxial dull or slightly glossy, pubescent or pilose, (hairs white, sometimes also ferruginous); proximal blade margins entire; juvenile blade yellowish green or reddish, glabrous, tomentose, or long-silky abaxially, hairs white, sometimes also ferruginous. |
Staminate flowers | adaxial nectary oblong or square, 0.3–0.7 mm; filaments distinct or basally connate, glabrous; anthers purple turning yellow, 0.4–0.6 mm. |
adaxial nectary narrowly oblong, oblong, or ovate, 0.4–0.9 mm; filaments distinct, glabrous or hairy on proximal 1/2 or basally; anthers yellow, shortly cylindrical or ovoid, 0.6–1 mm. |
Pistillate flowers | adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe; stipe 0.4–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm. |
adaxial nectary oblong, square, or obovate, 0.4–0.9 mm, shorter than stipe; stipe 1.2–2.7 mm; ovary pyriform or obclavate, tomentose or short-silky, beak slightly bulged below styles; ovules 12 per ovary; styles 0.2–0.5 mm; stigmas broadly cylindrical, 0.23–0.43–0.63 mm. |
Capsules | 3.2–8 mm. |
5–7 mm. |
Catkins | flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm; floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy. |
flowering before leaves emerge; staminate stout, 11–16 mm, flowering branchlet 0–5 mm; pistillate densely to loosely flowered, stout, 11–18 mm, flowering branchlet 0–3 mm; floral bract brown, black, or bicolor, 1–3 mm, apex acute, convex, or rounded, abaxially hairy, hairs straight or wavy. |
2n | = 38. |
= 76. |
Salix hastata |
Salix atrocinerea |
|
Phenology | Flowering early Jun-late Jul. | Flowering mid Mar-mid May. |
Habitat | Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests | Wooded wetlands, marshes, sandy beaches, mesic prairies, edges of birch-maple or oak woodlands |
Elevation | 0-1200 m (0-3900 ft) | 0-700 m (0-2300 ft) |
Distribution |
AK; NT; YT; Eurasia (Norway, Russia, Siberia) |
CT; MA; ME; MO; NC; NE; NJ; NY; PA; RI; WI; ON; Europe [Introduced in North America] |
Discussion | Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae. Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999). Hybrids: Salix hastata forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Hairs on abaxial leaf surfaces of Salix atrocinerea often are spreading to erect, and curly as in S. humilis. Usually, S. atrocinerea can be recognized by its closely spaced parallel tertiary venation. See 76. S. discolor and 81. S. cinerea for further comparative descriptions. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 133. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Vetrix > sect. Cinerella |
Sibling taxa | ||
Synonyms | S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei | S. cinerea var. atrocinerea, S. cinerea subsp. oleifolia |
Name authority | Linnaeus: Sp. Pl. 2: 1017. (1753) | Brotero: Fl. Lusit. 1: 31. (1804) |
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