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halberd willow

peach-leaf willow

Habit Plants 0.2–4 m. Stems: branches reddish brown, not glaucous, (slightly glossy), pilose; branchlets yellow-brown or red-brown, villous or pilose. Trees, 4–20 m. Stems: branches flexible to ± brittle at base, yellow to gray-brown, glabrous; branchlets yellow-brown, gray-brown, or red-brown, glabrous.
Leaves

stipules foliaceous, apex acute to acuminate;

petiole convex to flat, or shallowly grooved adaxially, 2–6(–9) mm, pilose or villous adaxially;

largest medial blade narrowly to broadly elliptic or narrowly ovate to ovate, 25–92 × 10–45 mm, 1.5–2.6(–3.4) times as long as wide, base convex, cuneate, or rounded, margins slightly revolute or flat, shallowly serrulate or entire, apex acuminate, acute, or convex, abaxial surface glaucous, sparsely pubescent, hairs wavy, adaxial dull to slightly glossy, pilose, sparsely pubescent or glabrous, midrib hairy, (hairs white and ferruginous);

proximal blade margins entire or finely serrulate;

juvenile blade sometimes reddish, sparsely pubescent abaxially, hairs white, sometimes also ferruginous.

stipules absent or rudimentary on early ones, foliaceous or rudimentary on late ones, apex rounded;

petiole (margins covering groove, not glandular or with spherical glands distally), 7–21 mm, glabrous or puberulent adaxially;

largest medial blade (sometimes amphistomatous), very narrowly elliptic, elliptic, lanceolate, or lorate, 55–130 × 24–37 mm, 2.8–6 times as long as wide, base convex, cuneate, or cordate, margins serrulate, apex acuminate to caudate, abaxial surface glaucous, glabrous, adaxial dull, glabrous or sparsely pubescent along midrib;

proximal blade margins entire or shallowly serrulate;

juvenile blade glabrous or pubescent abaxially, hairs white and/or ferruginous.

Staminate flowers

adaxial nectary oblong or square, 0.3–0.7 mm;

filaments distinct or basally connate, glabrous;

anthers purple turning yellow, 0.4–0.6 mm.

abaxial nectary 0.2–0.7 mm, adaxial nectary narrowly oblong to square, 0.3–0.8 mm, nectaries distinct;

stamens 3–7;

filaments hairy on proximal 1/2 or basally;

anthers 0.5–0.6 mm.

Pistillate flowers

adaxial nectary square or obovate, 0.3–0.6 mm, usually shorter than stipe;

stipe 0.4–1.2 mm;

ovary pyriform, glabrous, beak gradually tapering to styles;

ovules 12–22 per ovary;

styles connate (sometimes distinct 1/2 their lengths), 0.2–0.5 mm;

stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, or 2 plump lobes, 0.2–0.32–0.44 mm.

adaxial nectary square, 0.1–0.6 mm;

stipe 1.4–3.2 mm;

ovary pyriform, beak slightly bulged below styles;

ovules 16–18 per ovary;

styles 0.2–0.4 mm;

stigmas 0.24–0.31–0.4 mm.

Capsules

3.2–8 mm.

3–7 mm.

Catkins

flowering as leaves emerge; staminate slender, stout, or subglobose, 14.5–34.5 × 8–12 mm, flowering branchlet 1–7 mm; pistillate moderately densely or loosely flowered, slender or stout, 21–59 × 6–16 mm, flowering branchlet 1.5–9 mm;

floral bract brown or bicolor, 1.2–1.8 mm, apex acute to rounded, abaxially glabrate to hairy, hairs straight to wavy.

staminate 23–80 × 5–12 mm, flowering branchlet 3–28 mm; pistillate 41–110(–127 in fruit) × 8–16 mm, flowering branchlet 17–35 mm;

floral bract 1.5–2.8 mm, apex acute to rounded, entire or toothed, abaxially sparsely to moderately densely hairy proximally, hairs wavy; pistillate bract deciduous after flowering.

2n

= 38.

= 38.

Salix hastata

Salix amygdaloides

Phenology Flowering early Jun-late Jul. Flowering early Apr–Jun.
Habitat Sandy and gravelly river bars and floodplains, lakeshores, sand dunes and blowouts, Dryas tundra, alpine sedge meadows, balsam poplar thickets, openings in upland spruce-willow forests Moist to mesic floodplains, shores of lakes on sandy, silty, or gravelly substrates, marshes, wet sand dune slacks
Elevation 0-1200 m (0-3900 ft) 60-2400 m (200-7900 ft)
Distribution
from FNA
AK; NT; YT; Eurasia (Norway, Russia, Siberia)
[BONAP county map]
from FNA
AZ; CO; IA; ID; IL; IN; KS; KY; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix hastata is characterized by branches pilose with short, curved hairs; juvenile and mature leaves with white or ferruginous hairs adaxially, glaucous or not abaxially; and ovaries often reddish. See 61. S. barclayi and 64. S. farriae.

Subspecies subintegrifolia, characterized by entire or subentire leaf margins, is recognized in Flora Nordica as the major northern Eurasian race of Salix hastata, and is described as occurring across Russia into northwestern North America (B. Jonsell and T. Karlsson 2000+, vol. 1). Because leaf marginal toothing is highly variable throughout the species range (E. Hultén 1967), the subspecies is not recognized here or in Russia (A. K. Skvortsov 1999).

Hybrids:

Salix hastata forms natural hybrids with S. barclayi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Presence of Salix amygdaloides in Massachusetts, New Hampshire, and Vermont has not been verified; its occurrence in those New England states was reported by H. A. Gleason and A. Cronquist (1991), and by M. L. Fernald (1950).

Hybrids:

Salix amygdaloides forms natural hybrids with S. gooddingii and S. nigra. Hybrids with S. caroliniana (N. M. Glatfelter 1898) and S. eriocephala (M. L. Fernald 1950) have been reported; no convincing specimens have been seen. Controlled pollination between S. amygdaloides and S. eriocephala, S. interior, and S. petiolaris set no seed; controlled pollination with S. lucida produced a few seeds; some seedlings suffered necrosis in the cotyledon stage (A. Mosseler 1990).

Salix amygdaloides × S. gooddingii (S. ×wrightii Andersson): This hybrid occurs throughout the Rio Grande Valley, Texas, and New Mexico (C. K. Schneider 1919; C. R. Ball 1961), and at Happy and Rio Frio, Texas, and Virgil Run, Arizona. The leaves are somewhat glaucous abaxially, as in S. amygdaloides, but they are linear to narrowly elliptic and branchlets are sparsely pubescent as in S. gooddingii.

Salix amygdaloides × S. nigra (S. ×glatfelteri C. K. Schneider) resembles S. amygdaloides in leaves somewhat glaucous abaxially, but usually linear or narrowly elliptic, as in S. nigra. The stipules are not as prominent as in S. nigra but are foliaceous on late leaves; it should be expected wherever the ranges of the two species overlap. The hybrid is common in Missouri, where N. M. Glatfelter (1894) estimated that ca. 40% of the populations were hybrids, and in Illinois (R. H. Mohlenbrock 1980; G. Wilhelm, pers. comm.). Putative hybrids occur also in Ontario. Narrow leaves are typical of juvenile plants of S. amygdaloides but even at that stage they tend to be broadest at the midpoint or toward the base rather than in a midzone as in S. nigra.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 116. FNA vol. 7, p. 37.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Protitea > sect. Humboldtianae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. farriae var. walpolei, S. hastata subsp. subintegrifolia, S. walpolei
Name authority Linnaeus: Sp. Pl. 2: 1017. (1753) Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 114. (1858)
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