Salix glauca var. acutifolia |
Salix glauca |
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grey-leaf willow, point grey-leaf willow |
glaucous willow, gray willow, gray-leaf willow, grey-leaf willow |
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Habit | Plants 0.3–6 m. Stems: branches red-brown or brownish, villous or pilose to glabrescent; branchlets densely villous to glabrescent. | Plants 0.2–6 m, not clonal. | ||||||||||||
Stems | erect or decumbent; branches brownish, yellow-brown, gray-brown, or red-brown, villous or pilose to glabrescent; branchlets yellow-brown or red-brown, sparsely to densely villous or tomentose to glabrescent. |
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Leaves | stipules marcescent, rudimentary or foliaceous on early ones, foliaceous on late ones, usually prominent, linear to lanceolate, 2–17 mm, apex acuminate; petiole 4–27 mm, pilose adaxially; largest medial blade oblanceolate or obovate to narrowly elliptic, 43–82 × 12–39 mm, 2.2–4.8 times as long as wide, base cuneate or convex, apex acuminate or convex, abaxial surface very densely to sparsely villous-tomentose or short-silky, hairs straight, wavy, or curved, adaxial long-silky or pilose to glabrescent; proximal blade margins entire; juvenile blade densely villous or long-silky. |
stipules (marcescent or not), foliaceous or rudimentary on early and late ones; petiole (usually deeply to shallowly grooved adaxially), 1–27 mm, (much longer than subtended bud); largest medial blade usually hypostomatous, sometimes hemiamphistomatous or amphistomatous, usually narrowly elliptic, elliptic, usually oblanceolate or obovate, sometimes narrowly oblong or obovate, 27–82 × 6–39 mm, 1.4–4.8 times as long as wide, base usually cuneate or convex, sometimes rounded, rarely subcordate, margins slightly revolute or flat, usually entire, apex acute, acuminate, convex, or rounded, abaxial surface densely villous or villous-silky, tomentose, short- or long-silky, or pilose, hairs usually wavy or straight, sometimes curved, adaxial usually slightly glossy, sometimes dull, moderately densely villous, pilose, or long-silky to glabrescent; proximal blade margins entire or serrulate; juvenile blade sparsely or densely villous, tomentose, or long-silky abaxially. |
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Staminate flowers | abaxial nectary 0.1–0.6 mm, adaxial nectary narrowly oblong, oblong, or square, 0.6–0.9 mm; filaments distinct, glabrous. |
abaxial nectary 0.1–1 mm, adaxial nectary narrowly oblong, oblong, square, ovate, or flask-shaped, 0.5–1.3 mm, nectaries distinct, or connate and cup-shaped; filaments distinct or slightly or partly connate, glabrous, or hairy on proximal 1/2; anthers 0.4–0.8 mm. |
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Pistillate flowers | adaxial nectary sometimes flask-shaped; stipe 0.5–2.8 mm; ovary obclavate or pyriform, densely tomentose, short-silky, or pubescent, beak gradually tapering to styles; ovules 12–22 per ovary; styles connate to distinct 1/2 their lengths, 0.3–1.4 mm; stigmas slenderly to broadly cylindrical, 0.4–0.59–0.8 mm. |
abaxial nectary absent, adaxial nectary usually narrowly oblong, oblong, or ovate, sometimes flask-shaped, 0.4–1.8 mm, shorter to longer than stipe; stipe 0.3–2.8 mm; ovary pyriform or obclavate, densely villous, tomentose, short-silky, or pubescent, beak usually gradually tapering to styles, sometimes gradually tapering to or slightly bulged below styles; ovules 6–22 per ovary; styles connate to distinct 1/2 their lengths or more, 0.3–1.6 mm; stigmas flat, abaxially non-papillate with rounded tip, or slenderly or broadly cylindrical, 0.2–0.8 mm. |
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Capsules | 5–9 mm. |
4.5–9 mm. |
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Catkins | staminate 19–45 × 9–17 mm, flowering branchlet 2–9 mm; pistillate slender to stout, 24–83 × 8–17 mm, flowering branchlet 3–37 mm; floral bract tawny, light brown, or bicolor, 1.5–2.5 mm, apex convex to rounded, hairs wavy. |
staminate 14–53 × 5–17 mm, flowering branchlet 1–25 mm; pistillate densely to sometimes loosely flowered, slender, stout, subglobose, or globose, 15–83 × 7–21 mm, flowering branchlet 2–37 mm; floral bract tawny, brown, bicolor, or greenish, 1–3.4 mm, apex convex or rounded, entire, abaxially hairy, hairs wavy, crinkled, or straight. |
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2n | = 76, 95, 114. |
= 76, 95, 114, 152. |
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Salix glauca var. acutifolia |
Salix glauca |
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Phenology | Flowering late May-early Jul. | |||||||||||||
Habitat | Wet to mesic thickets, black spruce treed bogs, white spruce woods, floodplains, fens, swamps, subarctic thickets, alpine tundra | |||||||||||||
Elevation | 0-1200 m (0-3900 ft) | |||||||||||||
Distribution |
AK; BC; NT; YT |
AK; CO; MT; NM; OR; UT; WA; WY; AB; BC; MB; NS; NT; NU; ON; QC; SK; YT; Eurasia (China [Altay Shan], Chukotka, Mongolia, Novaya Zemlya, Russian Far East, Sakhalin, Scandinavia, arctic, e, w Siberia)
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Discussion | Hybrids: Variety acutifolia forms natural hybrids with Salix arctica and S. niphoclada. Variety acutifolia × Salix niphoclada, known from Alaska and Yukon, is intermediate between the parents. It combines the petioles, the narrowly elliptic-oblanceolate leaves with acute-attenuate apices, the stipules, and the narrow, loosely flowered catkins of S. niphoclada, and the relatively long petioles, the larger oblanceolate leaves, the larger stipules, and the densely flowered and broadly cylindrical catkins of S. glauca (G. W. Argus 1965). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 5 (4 in the flora). Occurrence of Salix glauca is disjunct in western Siberia; var. glauca is known from Scandinavia. Salix glauca is ubiquitous, highly polymorphic, and polyploid. It appears to have four major variations that are treated in different ways. European floras have treated it as an undivided polymorphic species(A. K. Skvortsov 1999), as several distinct species (K. H. Rechinger 1964b), or as comprising several subspecies (Rechinger 1993; G. W. Argus et al. 1999; B. Jonsell and T. Karlsson 2000+, vol. 1). In North America, Argus (1965) treated the four central tendencies as widely intergrading “phases” of a single species, later adopting varietal rank, whereas E. Hultén (1968) recognized them as subspecies. In view of the fact that the four elements are confluent over wide areas, there may be some merit in recognizing them as informal phases, but varietal rank is used here. All specimens from Iceland named Salix glauca are S. arctica and those from Svalbard, Norway, are S. lanata Linnaeus. The major reason for the high variability within Salix glauca seems to be high, and probably recurrent, polyploidy. Tetraploids, pentaploids, hexaploids, and octoploids are known in the species, and two of the subspecies include three ploidal levels. There also are intergrading geographical variations that are the basis for the recognition of infraspecific taxa. Hybrids: Salix glauca forms natural hybrids with S. arctica, S. arctophila, S. ballii, S. barclayi, S. boothii, S. brachycarpa, S. eastwoodiae, S. myricoides, S. niphoclada, S. pedicellaris, and S. planifolia. The following key will help identify the varieties, but there are extensive areas of overlap among them and many intermediates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 91. | FNA vol. 7, p. 89. | ||||||||||||
Parent taxa | ||||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | S. villosa var. acutifolia, S. desertorum, S. glauca subsp. acutifolia, S. glauca var. alicea, S. glauca subsp. desertorum, S. glauca var. perstipula, S. glauca var. poliophylla, S. glauca var. sericea | |||||||||||||
Name authority | (Hooker) C. K. Schneider: Bot. Gaz. 66: 327. (1918) | Linnaeus: Sp. Pl. 2: 1019. (1753) | ||||||||||||
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