Salix geyeriana |
Salix uva-ursi |
|
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Geyer willow, Geyer's willow |
bearberry willow |
|
Habit | Plants 0.6–5 m, (sometimes forming clones by stem fragmentation). | Plants 0.01–0.05 m, (dwarf), forming clonal compact mats by layering. |
Stems | branches (sometimes ± brittle at base), yellow-green, gray-brown, red-brown, or violet, usually glaucous, glabrous or sparsely tomentose; branchlets yellowish, yellow-brown, red-brown, or violet, (strongly glaucous or not), glabrous or sparsely to moderately densely pubescent, (buds caprea-type). |
prostrate, short-trailing; branches red-brown, gray-brown, or yellow-brown, glabrous; branchlets yellow-green or yellow-brown, glabrous or puberulent. |
Leaves | stipules usually absent or rudimentary (rarely foliaceous); petiole convex to flat, or shallowly to deeply grooved adaxially, 2–9 mm, velvety, short-silky, or pubescent adaxially; largest medial blade lorate, narrowly elliptic, or linear, 32–89 × 5.5–14 mm, 3.6–8.4(–11.3) times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire or distantly and shallowly serrulate, apex acute to acuminate, abaxial surface glaucous, glabrous or densely short- or long-silky, hairs (white, sometimes also ferruginous), straight, adaxial slightly glossy, densely short- or long-silky, especially midrib, to glabrescent, (hairs white, sometimes also ferruginous); proximal blade margins entire or distantly and shallowly serrulate; juvenile blade reddish or yellowish green, densely to sparsely long- or short-silky abaxially, hairs white, sometimes also ferruginous. |
stipules (sometimes marcescent) absent, rudimentary, or foliaceous on early ones, foliaceous on late ones; petiole (shallowly to deeply grooved adaxially), 2–6.5 mm; largest medial blade (marcescent), amphistomatous or hypostomatous, ovate, broadly obovate, oblanceolate, or elliptic, 4–23 × 3.5–10 mm, 1.7–3.6 times as long as wide, base convex or cuneate, margins flat, serrulate or crenulate, apex convex, acuminate, acute, or retuse, abaxial surface glaucous, usually glabrous (rarely few hairs), adaxial slightly or highly glossy, usually glabrous (rarely a few hairs); proximal blade margins entire or serrate; juvenile blade glabrous, pilose, or puberulent abaxially. |
Staminate flowers | adaxial nectary oblong, ovate, or square, 0.3–0.8(–0.9) mm; filaments distinct, glabrous or hairy on proximal 1/2 or basally; anthers yellow or purple turning yellow, ellipsoid or globose, 0.4–0.5(–0.6) mm. |
abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.4–0.9 mm; filaments distinct, glabrous; stamens usually 1, rarely 2; anthers ellipsoid or shortly cylindrical, 0.4–0.7 mm. |
Pistillate flowers | adaxial nectary oblong, square, or ovate, 0.2–1 mm; stipe (0.4–)1–2.8 mm; ovary pyriform (gourd-shaped), beak gradually tapering to styles; ovules 6–12 per ovary; styles 0.1–0.2(–0.6) mm; stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.2–0.3–0.44 mm. |
abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.5–0.8 mm, shorter to longer than stipe; stipe 0.3–1.6 mm; ovary ovoid or pyriform, glabrous, beak gradually tapering to styles; ovules 4–9 per ovary; styles 0.4–1 mm; stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.1–0.23–0.4 mm. |
Capsules | (3–)4–6 mm. |
3–5 mm. |
Catkins | flowering (before or) as leaves emerge; staminate globose, (1–1.1–)11–18 × 6–11 mm, flowering branchlet 1–5 mm; pistillate densely to loosely flowered, subglobose to globose, 8–21 × 7–17 mm, flowering branchlet 0.5–8 mm; floral bract tawny or brown (black), 1.2–2.8 mm, apex rounded or acute, abaxially hairy, hairs short, wavy or straight. |
staminate 9–19 × 5–8 mm, flowering branchlet 0.5–9 mm; pistillate densely flowered, slender to subglobose, 11–47(–55 in fruit) × 6–10 mm, flowering branchlet 2–10 mm; floral bract brown, black, tawny, light rose, or bicolor, 1.1–1.8 mm, apex rounded or acute, entire, abaxially sparsely hairy, hairs straight or wavy. |
2n | = 38. |
= 38. |
Salix geyeriana |
Salix uva-ursi |
|
Phenology | Flowering late Apr-late Jun. | Flowering mid Jun-early Aug. |
Habitat | Lowland wet streamsides, lakeshores, sedge meadows, springs, seepages, swamps, cienegas, fine-textured substrates | Exposed, often dry or moist, calcareous, serpentine, dioritic, and granitic rocks, boulders, gravel, sand on beaches, outcrops, in snowbeds |
Elevation | 10-3300 m (0-10800 ft) | 10-1200 m (0-3900 ft) |
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
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ME; NH; NY; VT; NL; NS; NU; QC; SPM; Greenland
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Discussion | Salix geyeriana is characterized by its dark gray appearance, slender, dark branches, narrow leaves long-silky on both surfaces, general absence of stipules, and small, subglobose catkins. Plants in the Pacific Northwest with foliaceous stipules may be hybrids or introgressants, but the other parent is unknown. Hybrids: Salix geyeriana forms natural hybrids with S. bebbiana, S. irrorata, S. lemmonii, S. ligulifolia, and S. pedicellaris. Alleged hybrids with S. sitchensis, based on plants from British Columbia with broader, more hairy leaves, and catkins longer than in S. geyeriana, but with the short stipes of S. sitchensis (J. K. Henry 1915), are unconvincing. Salix geyeriana × S. irrorata: A series of specimens from Arizona usually resemble S. geyeriana but have some characters of S. irrorata: foliaceous stipules, toothed leaf margins, catkins flowering before leaves emerge, shorter stipes (0.4–1.2 mm), and longer styles (0.2–0.6 mm). They also have some unique characters: proximal leaves sometimes serrulate, leaves sometimes amphistomatous, adaxial leaf surfaces mostly with ferruginous hairs, and ovaries sometimes gourd-shaped. Salix geyeriana × S. lemmonii is uncommon but in mixed stands of the parental species some plants resemble S. geyeriana in having relatively short, subspherical catkins, small anthers, and petioles sometimes with petiolar glands; and S. lemmonii in having leaf blades amphistomatous, margins serrulate, and foliaceous stipules on early leaves. Because the species have different chromosome numbers, hybrids may be infertile, but occasional seeds have been seen. This hybrid is known from California (Lassen and Sierra counties), Oregon (Jefferson and Lane counties), and near Victoria, British Columbia. Salix geyeriana × S. ligulifolia: Plants in Arizona with rudimentary stipules and leaves with ferruginous hairs may be this hybrid. Salix geyeriana × S. pedicellaris occurs in Washington. It has the white and ferruginous hairs on leaves and ovaries of S. geyeriana, and leaves glaucous adaxially with prominent 2 and 3 veins of S. pedicellaris. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Hybrids: Salix uva-ursi forms natural hybrids with S. herbacea. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 153. | FNA vol. 7, p. 73. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Geyerianae | Salicaceae > Salix > subg. Chamaetia > sect. Myrtosalix |
Sibling taxa | ||
Synonyms | S. geyeriana var. argentea, S. geyeriana var. meleina | S. ivigtutiana, S. myrsinites var. parvifolia |
Name authority | Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 122. (1858) | Pursh: Fl. Amer. Sept. 2: 610. (1813) |
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