Salix geyeriana |
Salix ovalifolia |
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Geyer willow, Geyer's willow |
arctic seashore or oval-leaf willow, oval-leaf willow |
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Habit | Plants 0.6–5 m, (sometimes forming clones by stem fragmentation). | Plants 0.02–0.05 m, not clonal or forming clones by layering. | ||||||||||||
Stems | branches (sometimes ± brittle at base), yellow-green, gray-brown, red-brown, or violet, usually glaucous, glabrous or sparsely tomentose; branchlets yellowish, yellow-brown, red-brown, or violet, (strongly glaucous or not), glabrous or sparsely to moderately densely pubescent, (buds caprea-type). |
trailing; branches yellow-brown, gray-brown, or red-brown, glabrous or hairy; branchlets yellow-green, yellow-brown, or red-brown, glabrous or pilose. |
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Leaves | stipules usually absent or rudimentary (rarely foliaceous); petiole convex to flat, or shallowly to deeply grooved adaxially, 2–9 mm, velvety, short-silky, or pubescent adaxially; largest medial blade lorate, narrowly elliptic, or linear, 32–89 × 5.5–14 mm, 3.6–8.4(–11.3) times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire or distantly and shallowly serrulate, apex acute to acuminate, abaxial surface glaucous, glabrous or densely short- or long-silky, hairs (white, sometimes also ferruginous), straight, adaxial slightly glossy, densely short- or long-silky, especially midrib, to glabrescent, (hairs white, sometimes also ferruginous); proximal blade margins entire or distantly and shallowly serrulate; juvenile blade reddish or yellowish green, densely to sparsely long- or short-silky abaxially, hairs white, sometimes also ferruginous. |
stipules usually absent or rudimentary, rarely foliaceous; petiole (deeply to shallowly grooved adaxially), 1.1–16 mm, (glabrous); largest medial blade hypostomatous, narrowly to broadly elliptic, circular, subcircular, or obovate, 13–46 × 7–20 mm, 1–3.4 times as long as wide, base subcordate, cordate, rounded, or convex, margins slightly revolute or flat, entire, sometimes ciliate, apex convex, rounded, acuminate, acute, or retuse, abaxial surface glabrous, villous, long-silky, pubescent, or pilose, hairs wavy or straight, adaxial highly glossy, usually glabrous; proximal blade margins entire; juvenile blade (reddish or yellowish green), pilose, villous, or long-silky abaxially. |
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Staminate flowers | adaxial nectary oblong, ovate, or square, 0.3–0.8(–0.9) mm; filaments distinct, glabrous or hairy on proximal 1/2 or basally; anthers yellow or purple turning yellow, ellipsoid or globose, 0.4–0.5(–0.6) mm. |
abaxial nectary 0.6–1 mm, adaxial nectary oblong or ovate, 0.6–1.6 mm, nectaries distinct or connate and cup-shaped; filaments distinct or connate less than 1/2 their lengths (glabrous); anthers elliptic, short-cylindrical, or globose, 0.3–0.5(–0.6) mm. |
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Pistillate flowers | adaxial nectary oblong, square, or ovate, 0.2–1 mm; stipe (0.4–)1–2.8 mm; ovary pyriform (gourd-shaped), beak gradually tapering to styles; ovules 6–12 per ovary; styles 0.1–0.2(–0.6) mm; stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.2–0.3–0.44 mm. |
abaxial nectary (0–)0.4–0.6 mm, adaxial nectary longer than stipe; stipe 0.2–1.4 mm; ovary obclavate or pyriform, glaucous or not, usually glabrous or tomentose, sometimes pubescent or villous, beak abruptly tapering to styles; ovules 10–15 per ovary; styles 0.2–0.8 mm; stigmas flat, abaxially non-papillate with pointed tip, or slenderly cylindrical, 0.32–0.41–0.64 mm. |
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Capsules | (3–)4–6 mm. |
5.2–9.6 mm. |
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Catkins | flowering (before or) as leaves emerge; staminate globose, (1–1.1–)11–18 × 6–11 mm, flowering branchlet 1–5 mm; pistillate densely to loosely flowered, subglobose to globose, 8–21 × 7–17 mm, flowering branchlet 0.5–8 mm; floral bract tawny or brown (black), 1.2–2.8 mm, apex rounded or acute, abaxially hairy, hairs short, wavy or straight. |
staminate 4.8–46 × 5–11 mm, flowering branchlet 1.5–24 mm; pistillate moderately densely flowered, stout, subglobose, globose, or slender, 6.3–50 × 5–28 mm, flowering branchlet 2.5–22 mm; floral bract brown, greenish, or bicolor, 1.2–2 mm, apex rounded, entire or, sometimes, 2-fid, abaxially hairy, hairs straight or wavy. |
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2n | = 38. |
= 38. |
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Salix geyeriana |
Salix ovalifolia |
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Phenology | Flowering late Apr-late Jun. | |||||||||||||
Habitat | Lowland wet streamsides, lakeshores, sedge meadows, springs, seepages, swamps, cienegas, fine-textured substrates | |||||||||||||
Elevation | 10-3300 m (0-10800 ft) | |||||||||||||
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
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AK; NT; YT; Asia |
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Discussion | Salix geyeriana is characterized by its dark gray appearance, slender, dark branches, narrow leaves long-silky on both surfaces, general absence of stipules, and small, subglobose catkins. Plants in the Pacific Northwest with foliaceous stipules may be hybrids or introgressants, but the other parent is unknown. Hybrids: Salix geyeriana forms natural hybrids with S. bebbiana, S. irrorata, S. lemmonii, S. ligulifolia, and S. pedicellaris. Alleged hybrids with S. sitchensis, based on plants from British Columbia with broader, more hairy leaves, and catkins longer than in S. geyeriana, but with the short stipes of S. sitchensis (J. K. Henry 1915), are unconvincing. Salix geyeriana × S. irrorata: A series of specimens from Arizona usually resemble S. geyeriana but have some characters of S. irrorata: foliaceous stipules, toothed leaf margins, catkins flowering before leaves emerge, shorter stipes (0.4–1.2 mm), and longer styles (0.2–0.6 mm). They also have some unique characters: proximal leaves sometimes serrulate, leaves sometimes amphistomatous, adaxial leaf surfaces mostly with ferruginous hairs, and ovaries sometimes gourd-shaped. Salix geyeriana × S. lemmonii is uncommon but in mixed stands of the parental species some plants resemble S. geyeriana in having relatively short, subspherical catkins, small anthers, and petioles sometimes with petiolar glands; and S. lemmonii in having leaf blades amphistomatous, margins serrulate, and foliaceous stipules on early leaves. Because the species have different chromosome numbers, hybrids may be infertile, but occasional seeds have been seen. This hybrid is known from California (Lassen and Sierra counties), Oregon (Jefferson and Lane counties), and near Victoria, British Columbia. Salix geyeriana × S. ligulifolia: Plants in Arizona with rudimentary stipules and leaves with ferruginous hairs may be this hybrid. Salix geyeriana × S. pedicellaris occurs in Washington. It has the white and ferruginous hairs on leaves and ovaries of S. geyeriana, and leaves glaucous adaxially with prominent 2 and 3 veins of S. pedicellaris. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). The varieties of Salix ovalifolia are relatively minor variants; their ranges overlap and their differences in leaf shape and ovary indumentum intergrade (G. W. Argus 1969, 1973). The only one with a more or less distinctive geographical distribution is var. cyclophylla; where its range overlaps with var. ovalifolia there is intergradation. Variety arctolitoralis, which is characterized by larger leaves and catkins, may be an ecotype. Variety glacialis is known only from near Point Barrow, Alaska. E. Hultén (1968) suggested, probably based on its often tomentose ovaries, that it is S. arctica × S. ovalifolia. All varieties of the species have some plants with hairy ovaries, but the suggestion that this character is an indication of hybridization deserves study. Hybrids: Salix ovalifolia forms natural hybrids with S. arctica and S. fuscescens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 153. | FNA vol. 7, p. 77. | ||||||||||||
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Geyerianae | Salicaceae > Salix > subg. Chamaetia > sect. Ovalifoliae | ||||||||||||
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Synonyms | S. geyeriana var. argentea, S. geyeriana var. meleina | |||||||||||||
Name authority | Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 122. (1858) | Trautvetter: Nouv. Mém. Soc. Imp. Naturalistes Moscou 2: 306, plate 13. (1832) | ||||||||||||
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