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Geyer willow, Geyer's willow

Jepson's willow

Habit Plants 0.6–5 m, (sometimes forming clones by stem fragmentation). Shrubs, 1–3 m, (sometimes forming clones by stem fragmentation).
Stems

branches (sometimes ± brittle at base), yellow-green, gray-brown, red-brown, or violet, usually glaucous, glabrous or sparsely tomentose;

branchlets yellowish, yellow-brown, red-brown, or violet, (strongly glaucous or not), glabrous or sparsely to moderately densely pubescent, (buds caprea-type).

branches (highly brittle at base), yellow-brown or red-brown, weakly glaucous or not, (dull or slightly glossy), velvety or short-silky to glabrescent;

branchlets gray-brown or red-brown, sparsely or moderately densely short-silky or velvety, (buds caprea-type or intermediate).

Leaves

stipules usually absent or rudimentary (rarely foliaceous);

petiole convex to flat, or shallowly to deeply grooved adaxially, 2–9 mm, velvety, short-silky, or pubescent adaxially;

largest medial blade lorate, narrowly elliptic, or linear, 32–89 × 5.5–14 mm, 3.6–8.4(–11.3) times as long as wide, base cuneate or convex, margins flat or slightly revolute, entire or distantly and shallowly serrulate, apex acute to acuminate, abaxial surface glaucous, glabrous or densely short- or long-silky, hairs (white, sometimes also ferruginous), straight, adaxial slightly glossy, densely short- or long-silky, especially midrib, to glabrescent, (hairs white, sometimes also ferruginous);

proximal blade margins entire or distantly and shallowly serrulate;

juvenile blade reddish or yellowish green, densely to sparsely long- or short-silky abaxially, hairs white, sometimes also ferruginous.

stipules usually rudimentary, sometimes foliaceous on early ones, foliaceous on late ones, apex acute;

petiole shallowly grooved, or convex to flat adaxially, 3–12 mm, short-silky or velvety adaxially;

largest medial blade lorate, narrowly oblanceolate, or oblanceolate, 43–74–103 × 8–25 mm, 2.9–4.8–7.7 times as long as wide, base convex or cuneate, margins slightly revolute to flat, entire, apex acuminate, convex, or acute, abaxial surface glaucous (sometimes obscured by hairs), densely short-silky, hairs straight, adaxial dull (sometimes thinly glaucous) to slightly glossy, sparsely short- or long-silky;

proximal blade margins entire;

juvenile blade yellowish green or reddish, densely long-silky abaxially, hairs white.

Staminate flowers

adaxial nectary oblong, ovate, or square, 0.3–0.8(–0.9) mm;

filaments distinct, glabrous or hairy on proximal 1/2 or basally;

anthers yellow or purple turning yellow, ellipsoid or globose, 0.4–0.5(–0.6) mm.

adaxial nectary narrowly oblong to oblong, 0.4–0.8 mm;

stamens 1 or 2;

filaments distinct or connate less than 1/2 their lengths, hairy on proximal 1/2;

anthers purple turning yellow, ellipsoid or shortly cylindrical, 0.6–0.8 mm.

Pistillate flowers

adaxial nectary oblong, square, or ovate, 0.2–1 mm;

stipe (0.4–)1–2.8 mm;

ovary pyriform (gourd-shaped), beak gradually tapering to styles;

ovules 6–12 per ovary;

styles 0.1–0.2(–0.6) mm;

stigmas flat, abaxially non-papillate with rounded or pointed tip, 0.2–0.3–0.44 mm.

adaxial nectary oblong to narrowly oblong or flask-shaped, 0.3–0.6 mm, shorter than or equal to stipe;

stipe 0.4–1.2 mm;

ovary pyriform, densely long-silky, beak gradually tapering to styles;

ovules 13–18 per ovary;

styles 0.4–0.6 mm;

stigmas flat, abaxially non-papillate with rounded to pointed tip, 0.2–0.3 mm.

Capsules

(3–)4–6 mm.

3–5 mm.

Catkins

flowering (before or) as leaves emerge; staminate globose, (1–1.1–)11–18 × 6–11 mm, flowering branchlet 1–5 mm; pistillate densely to loosely flowered, subglobose to globose, 8–21 × 7–17 mm, flowering branchlet 0.5–8 mm;

floral bract tawny or brown (black), 1.2–2.8 mm, apex rounded or acute, abaxially hairy, hairs short, wavy or straight.

flowering as leaves emerge; staminate stout or subglobose, 16–16.5 × 11–13 mm, flowering branchlet 2.5–3 mm; pistillate moderately densely flowered, slender to stout, 33–55 × 10–11 mm, flowering branchlet 1.5–7 mm;

floral bract brown or tawny, 0.8–2 mm, apex rounded or acute, abaxially hairy, hairs straight.

2n

= 38.

Salix geyeriana

Salix jepsonii

Phenology Flowering late Apr-late Jun. Flowering mid-late Jun.
Habitat Lowland wet streamsides, lakeshores, sedge meadows, springs, seepages, swamps, cienegas, fine-textured substrates Margins of lakes and streams, wet meadows, gravel, rocky, or bouldery substrates, granite
Elevation 10-3300 m (0-10800 ft) 1000-3400 m (3300-11200 ft)
Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix geyeriana is characterized by its dark gray appearance, slender, dark branches, narrow leaves long-silky on both surfaces, general absence of stipules, and small, subglobose catkins. Plants in the Pacific Northwest with foliaceous stipules may be hybrids or introgressants, but the other parent is unknown.

Hybrids:

Salix geyeriana forms natural hybrids with S. bebbiana, S. irrorata, S. lemmonii, S. ligulifolia, and S. pedicellaris. Alleged hybrids with S. sitchensis, based on plants from British Columbia with broader, more hairy leaves, and catkins longer than in S. geyeriana, but with the short stipes of S. sitchensis (J. K. Henry 1915), are unconvincing.

Salix geyeriana × S. irrorata: A series of specimens from Arizona usually resemble S. geyeriana but have some characters of S. irrorata: foliaceous stipules, toothed leaf margins, catkins flowering before leaves emerge, shorter stipes (0.4–1.2 mm), and longer styles (0.2–0.6 mm). They also have some unique characters: proximal leaves sometimes serrulate, leaves sometimes amphistomatous, adaxial leaf surfaces mostly with ferruginous hairs, and ovaries sometimes gourd-shaped.

Salix geyeriana × S. lemmonii is uncommon but in mixed stands of the parental species some plants resemble S. geyeriana in having relatively short, subspherical catkins, small anthers, and petioles sometimes with petiolar glands; and S. lemmonii in having leaf blades amphistomatous, margins serrulate, and foliaceous stipules on early leaves. Because the species have different chromosome numbers, hybrids may be infertile, but occasional seeds have been seen. This hybrid is known from California (Lassen and Sierra counties), Oregon (Jefferson and Lane counties), and near Victoria, British Columbia.

Salix geyeriana × S. ligulifolia: Plants in Arizona with rudimentary stipules and leaves with ferruginous hairs may be this hybrid.

Salix geyeriana × S. pedicellaris occurs in Washington. It has the white and ferruginous hairs on leaves and ovaries of S. geyeriana, and leaves glaucous adaxially with prominent 2 and 3 veins of S. pedicellaris.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

R. D. Dorn (2000) justified treating Salix jepsonii as S. sitchensis var. angustifolia based on specimens of S. jepsonii from southern Oregon. Salix jepsonii does occur in southern Oregon but is distinct enough from S. sitchensis to be treated as a species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 153. FNA vol. 7, p. 160.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Geyerianae Salicaceae > Salix > subg. Vetrix > sect. Sitchenses
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. geyeriana var. argentea, S. geyeriana var. meleina S. sitchensis var. angustifolia, S. pellita var. angustifolia, S. sitchensis var. ralphiana
Name authority Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 122. (1858) C. K. Schneider: J. Arnold Arbor. 1: 89. (1919)
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