Salix farriae |
Salix jepsonii |
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Farr's willow |
Jepson's willow |
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Habit | Plants 0.2–1.5(–2) m. Stems: branches red-brown, not glaucous to strongly glaucous on buds, glabrous or puberulent at nodes; branchlets yellow-brown or red-brown, (sometimes weakly glaucous), glabrous or puberulent, (inner membranaceous bud-scale layer free, separating from outer layer). | Shrubs, 1–3 m, (sometimes forming clones by stem fragmentation). |
Stems | branches (highly brittle at base), yellow-brown or red-brown, weakly glaucous or not, (dull or slightly glossy), velvety or short-silky to glabrescent; branchlets gray-brown or red-brown, sparsely or moderately densely short-silky or velvety, (buds caprea-type or intermediate). |
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Leaves | stipules absent, rudimentary or foliaceous on early ones, foliaceous on late ones, apex acute; petiole shallowly grooved, or convex to flat adaxially, 5–8 mm, puberulent adaxially; largest medial blade narrowly elliptic or elliptic, (20–)30–65(–75) × (8–)10–30(–35) mm, 1.8–3.7 times as long as wide, base convex, rounded, or cuneate, margins slightly revolute or flat, entire or shallowly serrulate, apex acute, acuminate, or convex, abaxial surface glaucous, glabrous or glabrescent, adaxial slightly glossy or dull, glabrous or pilose, midrib sparsely pubescent, hairs short, white, and ferruginous; proximal blade margins entire or serrulate; juvenile blade green, glabrous, or midrib sparsely villous abaxially, hairs usually white and ferruginous. |
stipules usually rudimentary, sometimes foliaceous on early ones, foliaceous on late ones, apex acute; petiole shallowly grooved, or convex to flat adaxially, 3–12 mm, short-silky or velvety adaxially; largest medial blade lorate, narrowly oblanceolate, or oblanceolate, 43–74–103 × 8–25 mm, 2.9–4.8–7.7 times as long as wide, base convex or cuneate, margins slightly revolute to flat, entire, apex acuminate, convex, or acute, abaxial surface glaucous (sometimes obscured by hairs), densely short-silky, hairs straight, adaxial dull (sometimes thinly glaucous) to slightly glossy, sparsely short- or long-silky; proximal blade margins entire; juvenile blade yellowish green or reddish, densely long-silky abaxially, hairs white. |
Staminate flowers | adaxial nectary oblong, square, or ovate, 0.2–0.9 mm; filaments distinct, glabrous; anthers yellow, 0.3–0.6 mm. |
adaxial nectary narrowly oblong to oblong, 0.4–0.8 mm; stamens 1 or 2; filaments distinct or connate less than 1/2 their lengths, hairy on proximal 1/2; anthers purple turning yellow, ellipsoid or shortly cylindrical, 0.6–0.8 mm. |
Pistillate flowers | adaxial nectary oblong or ovate, 0.4–0.8 mm, shorter than stipe; stipe 0.5–1.2 mm; ovary pyriform, glabrous, beak gradually tapering to styles; ovules 12–19 per ovary; styles 0.3–1.2 mm; stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.2–0.3–0.56 mm. |
adaxial nectary oblong to narrowly oblong or flask-shaped, 0.3–0.6 mm, shorter than or equal to stipe; stipe 0.4–1.2 mm; ovary pyriform, densely long-silky, beak gradually tapering to styles; ovules 13–18 per ovary; styles 0.4–0.6 mm; stigmas flat, abaxially non-papillate with rounded to pointed tip, 0.2–0.3 mm. |
Capsules | 3–7 mm. |
3–5 mm. |
Catkins | flowering as leaves emerge; staminate stout, 11–25 × 6–11 mm, flowering branchlet 1–5 mm; pistillate densely or loosely flowered, stout, 14–38.5 × 8–14 mm, flowering branchlet 1.5–14 mm; floral bract brown, black, or bicolor, 0.7–2 mm, apex rounded to convex, abaxially hairy, hairs wavy. |
flowering as leaves emerge; staminate stout or subglobose, 16–16.5 × 11–13 mm, flowering branchlet 2.5–3 mm; pistillate moderately densely flowered, slender to stout, 33–55 × 10–11 mm, flowering branchlet 1.5–7 mm; floral bract brown or tawny, 0.8–2 mm, apex rounded or acute, abaxially hairy, hairs straight. |
Salix farriae |
Salix jepsonii |
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Phenology | Flowering late May-late Jul. | Flowering mid-late Jun. |
Habitat | Wet montane to subalpine meadows, stream banks | Margins of lakes and streams, wet meadows, gravel, rocky, or bouldery substrates, granite |
Elevation | 600-2700 m (2000-8900 ft) | 1000-3400 m (3300-11200 ft) |
Distribution |
ID; MT; OR; WY; AB; BC; NT; YT
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CA; NV; OR
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Discussion | Salix farriae is a cordilleran species ranging from Wyoming to central British Columbia with disjunct occurrences in northwestern British Columbia, western Northwest Territories, and southern Yukon. It is related to S. hastata, an amphiberingian species ranging from Scandinavia to southwestern Yukon and northwestern Northwest Territories. There may be reasons for treating these slightly different plants as S. hastata var. farriae, but R. D. Dorn (1975) maintained them as a species based on flavonoid differences. In a phenetic study (G. W. Argus 2007), the two taxa had dissimilarity values at the same level as other closely related species. They are treated here as species, primarily because their ranges are disjunct. They can be separated as follows: Salix farriae is distinguished from S. hastata by having largest medial blades narrowly elliptic to elliptic, pistillate nectaries oblong or ovate, stipules on early leaves absent or rudimentary (sometimes foliaceous), branches strongly to weakly glaucous or not, floral bract apices rounded, and plants of the cordillera in Alberta and British Columbia, in Idaho, Montana, Oregon, and Wyoming; S. hastata has largest medial blades narrowly elliptic to broadly elliptic or broadly obovate, pistillate nectaries square, stipules on early leaves foliaceous (sometimes rudimentary), branches not glaucous, floral bract apices acute or rounded, and plants of Alaska, Northwest Territories, and Yukon. Salix farriae and S. barclayi are sympatric in western Canada and the Pacific Northwest, where they are difficult to separate. Salix farriae can often be recognized by its largest medial leaves with at least some minute, ferruginous hairs on the adaxial midrib or blade surfaces; ferruginous hairs do not occur in S. barclayi. Its leaf margins also tend to be more nearly entire, but relatively short teeth are not infrequent. Such plants are sometimes interpreted as intergrades between S. farriae and S. barclayi (R. D. Dorn 1975). The variable leaf toothing also occurs in S. hastata and may not be a reliable indicator of intergradation. Salix farriae also differs from S. barclayi in usually having shorter anthers, 0.3–0.6 mm versus 0.6–1 mm in S. barclayi. See 61. S. barclayi. Hybrids: Salix farriae forms natural hybrids with S. barclayi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. D. Dorn (2000) justified treating Salix jepsonii as S. sitchensis var. angustifolia based on specimens of S. jepsonii from southern Oregon. Salix jepsonii does occur in southern Oregon but is distinct enough from S. sitchensis to be treated as a species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 116. | FNA vol. 7, p. 160. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Hastatae | Salicaceae > Salix > subg. Vetrix > sect. Sitchenses |
Sibling taxa | ||
Synonyms | S. farriae var. microserrulata, S. hastata var. farriae | S. sitchensis var. angustifolia, S. pellita var. angustifolia, S. sitchensis var. ralphiana |
Name authority | C. R. Ball: Contr. U.S. Natl. Herb. 22: 321. (1921) | C. K. Schneider: J. Arnold Arbor. 1: 89. (1919) |
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