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furry willow, heart-leaf or sand dune willow, heartleaf willow, sand dune willow

bearberry willow

Habit Plants 0.4–3 m, (often forming clones by layering or stem fragmentation). Plants 0.01–0.05 m, (dwarf), forming clonal compact mats by layering.
Stems

branches (sometimes ± brittle at base), red-brown, not glaucous (slightly glossy), tomentose to glabrescent;

branchlets red-brown, moderately to very densely villous.

prostrate, short-trailing;

branches red-brown, gray-brown, or yellow-brown, glabrous;

branchlets yellow-green or yellow-brown, glabrous or puberulent.

Leaves

stipules foliaceous, apex acute or rounded;

petiole shallowly grooved adaxially, 1–13 mm, tomentose adaxially;

largest medial blade narrowly oblong, or narrowly to broadly elliptic, 33–88 × 13–45 mm, 1.6–3.2 times as long as wide, base cordate, rounded, or convex, margins flat or slightly revolute, serrulate or spinulose-serrulate, apex acuminate, abaxial surface not glaucous, moderately densely villous to glabrescent, midrib remaining hairy, hairs straight or wavy, adaxial dull or slightly glossy, very densely villous to glabrescent, midrib remaining hairy;

proximal blade margins entire or serrulate;

juvenile blade yellowish green, abaxially villous or midrib long-silky, hairs white.

stipules (sometimes marcescent) absent, rudimentary, or foliaceous on early ones, foliaceous on late ones;

petiole (shallowly to deeply grooved adaxially), 2–6.5 mm;

largest medial blade (marcescent), amphistomatous or hypostomatous, ovate, broadly obovate, oblanceolate, or elliptic, 4–23 × 3.5–10 mm, 1.7–3.6 times as long as wide, base convex or cuneate, margins flat, serrulate or crenulate, apex convex, acuminate, acute, or retuse, abaxial surface glaucous, usually glabrous (rarely few hairs), adaxial slightly or highly glossy, usually glabrous (rarely a few hairs);

proximal blade margins entire or serrate;

juvenile blade glabrous, pilose, or puberulent abaxially.

Staminate flowers

adaxial nectary oblong or narrowly oblong, 0.3–1.3 mm;

filaments distinct, glabrous;

anthers yellow, (ellipsoid or shortly cylindrical), 0.6–0.8 mm.

abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.4–0.9 mm;

filaments distinct, glabrous;

stamens usually 1, rarely 2;

anthers ellipsoid or shortly cylindrical, 0.4–0.7 mm.

Pistillate flowers

adaxial nectary oblong or narrowly oblong, 0.4–1.3 mm, about same length as stipe;

stipe 0.5–1.4 mm;

ovary pyriform or obclavate, glabrous, beak slightly bulged below styles;

ovules 11–24 per ovary;

styles 0.7–1.6 mm;

stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.2–0.3–0.36 mm.

abaxial nectary absent, adaxial nectary narrowly oblong or oblong, 0.5–0.8 mm, shorter to longer than stipe;

stipe 0.3–1.6 mm;

ovary ovoid or pyriform, glabrous, beak gradually tapering to styles;

ovules 4–9 per ovary;

styles 0.4–1 mm;

stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.1–0.23–0.4 mm.

Capsules

3.6–7 mm.

3–5 mm.

Catkins

flowering as leaves emerge; staminate stout, 17–40 × 8–15 mm, flowering branchlet 1–8 mm; pistillate moderately densely flowered, stout or slender, 27–65 × 8–19 mm, flowering branchlet 3–16 mm;

floral bract brown, 1–2.6 mm, apex acute or rounded, abaxially hairy, hairs straight or wavy.

staminate 9–19 × 5–8 mm, flowering branchlet 0.5–9 mm; pistillate densely flowered, slender to subglobose, 11–47(–55 in fruit) × 6–10 mm, flowering branchlet 2–10 mm;

floral bract brown, black, tawny, light rose, or bicolor, 1.1–1.8 mm, apex rounded or acute, entire, abaxially sparsely hairy, hairs straight or wavy.

2n

= 38.

= 38.

Salix cordata

Salix uva-ursi

Phenology Flowering mid Apr-early Jul. Flowering mid Jun-early Aug.
Habitat Sand dunes and beaches Exposed, often dry or moist, calcareous, serpentine, dioritic, and granitic rocks, boulders, gravel, sand on beaches, outcrops, in snowbeds
Elevation 0-200 m (0-700 ft) 10-1200 m (0-3900 ft)
Distribution
from FNA
IL; IN; MI; NY; PA; WI; NL; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
ME; NH; NY; VT; NL; NS; NU; QC; SPM; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix cordata seems to be rare in Labrador, Newfoundland, and Quebec; the species needs further study in those areas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybrids:

Salix uva-ursi forms natural hybrids with S. herbacea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 106. FNA vol. 7, p. 73.
Parent taxa Salicaceae > Salix > subg. Vetrix > sect. Hastatae Salicaceae > Salix > subg. Chamaetia > sect. Myrtosalix
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. adenophylla, S. syrticola S. ivigtutiana, S. myrsinites var. parvifolia
Name authority Michaux: Fl. Bor.-Amer. 2: 225. (1803) Pursh: Fl. Amer. Sept. 2: 610. (1813)
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