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Cascade willow, Cascades willow

firmleaf willow, tall blueberry willow

Habit Plants 0.03–0.1 m, forming clones by rhizomes. Plants 1–7 m. Stems: branches gray-brown, red-brown, or yellow-brown, glaucous to strongly so (slightly to highly glossy), villous to glabrescent; branchlets gray-brown, red-brown, yellow-brown, or yellow-green, (not or weakly glaucous), pilose, densely villous, or tomentose.
Stems

erect or trailing;

branches yellow-brown or gray-brown, (sometimes weakly glaucous), glabrous;

branchlets yellow-green or yellow-brown, glabrous or puberulent.

Leaves

(marcescent);

stipules absent;

petiole 1.5–5 mm;

largest medial blade usually amphistomatous, narrowly elliptic or elliptic, 9–26 × 3.8–7.5 mm, 2.4–4.3 times as long as wide, base cuneate, margins flat, entire, ciliate, apex acute, acuminate, or convex, abaxial surface glabrous or pilose, hairs wavy, adaxial slightly glossy, glabrous or pilose;

proximal blade margins entire;

juvenile blade glabrous or sparsely villous abaxially.

stipules rudimentary or foliaceous on early ones, foliaceous on late ones, (0.6–8 mm), apex acute or obtuse;

petiole shallowly to deeply grooved adaxially, 2.5–8 mm, glabrous or villous adaxially;

largest medial blade (sometimes amphistomatous), narrowly to broadly elliptic, oblong to oblanceolate or obovate, 32–109 × 10–47 mm, 1.8–4.8 times as long as wide, base convex, cuneate, or subcordate, sometimes cordate, margins flat, entire, crenate, or serrulate, apex acute, convex, or acuminate, abaxial surface not glaucous, glabrous or pilose, hairs (white, sometimes also ferruginous), wavy, adaxial slightly glossy, glabrous, pilose, pubescent, moderately densely short-silky, or velvety, midrib remaining pilose or short-hairy, (hairs sometimes also ferruginous, straight and geniculate);

proximal blade margins serrulate or entire;

juvenile blade sometimes reddish or yellowish green, abaxially glabrous, or midrib sparsely pubescent, or densely villous or short-silky, hairs white, sometimes also ferruginous.

Staminate flowers

abaxial nectary (0–)0.2–0.6 mm, adaxial nectary oblong or square, 0.4–1.2 mm, nectaries distinct;

filaments distinct or basally connate;

anthers ellipsoid, 0.3–0.6 mm.

adaxial nectary oblong square, 0.2–0.4–0.6 mm;

filaments distinct, glabrous;

anthers purple turning yellow, 0.4–0.7 mm.

Pistillate flowers

adaxial nectary oblong or square, 0.5–1.2 mm, longer than stipe;

stipe 0–0.6 m;

ovary pyriform, usually densely villous, beak abruptly tapering to or slightly bulged below styles;

ovules 6–10 per ovary;

styles 0.3–1 mm;

stigmas slenderly cylindrical, 0.28–0.39–0.56 mm.

adaxial nectary square or oblong, 0.2–0.4 mm, shorter than stipe;

stipe 0.8–1.4 mm;

ovary pyriform, glabrous, beak slightly bulged below styles;

ovules 11–18 per ovary;

styles 0.4–1.6 mm;

stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.16–0.24–0.32 mm.

Capsules

3.5–5 mm.

4.4–6.4 mm.

Catkins

staminate (20–50 flowers), 12.5–26.5 × 5.5–9 mm, flowering branchlet 1–11 mm; pistillate moderately densely to loosely flowered, (15–35(–43) flowers), stout or subglobose, 10–23(–30 in fruit) × 5–8 mm, flowering branchlet 2–9 mm;

floral bract brown, 1.6–2.6 mm, apex rounded, entire, abaxially sparsely hairy or ciliate, hairs wavy or straight.

flowering as leaves emerge; staminate stout, 16.5–35.5 × 7–15 mm, flowering branchlet 0.5–12 mm; pistillate moderately or densely flowered, slender or stout, 10.5–68 × 5–20 mm, flowering branchlet 0.5–10 mm;

floral bract brown, black, tawny, or bicolor, 0.6–1.1 mm, apex retuse, abaxially hairy, hairs long-wavy or curly.

2n

= 76.

Salix cascadensis

Salix pseudomyrsinites

Phenology Flowering early Jul-early Aug. Flowering early May-early Jul.
Habitat Mesic to dry rocky slopes, ridges, high subalpine and alpine tundra Shores of lakes and streams, dwarf-birch thickets, fens, marl bogs, rarely in treed bogs
Elevation 2200-3900 m (7200-12800 ft) 40-1000 m (100-3300 ft)
Distribution
from FNA
CO; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AB; BC; MB; NT; NU; ON; SK; YT
Discussion

The morphological variability of Salix cascadensis is not well understood. Typically, it has leaves that are narrow, sharply pointed, and glaucous abaxially, catkins 15–43-flowered, dark brown floral bracts, and ovaries very densely hairy. Specimens with leaves not glaucous abaxially, catkins relatively few-flowered, and ovaries either sparsely hairy throughout, hairy only on beaks, or hairy in streaks, may be hybrids but the glabrous ovaried S. cascadensis var. thompsonii shows no obvious signs of hybridization.

Hybrids:

Salix cascadensis is suspected to hybridize with S. barclayi.

Salix cascadensis × S. rotundifolia var. dodgeana resembles the former in its narrow, acute to acuminate leaves and the latter in leaves not glaucous abaxially, tawny floral bracts, relatively short catkins, and glabrous ovaries. The catkins are intermediate in being 6–7-flowered.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Salix pseudomyrsinites occurs in Nunavut on Akimiski Island in James Bay.

Salix pseudomyrsinites and S. myrtillifolia, although sometimes treated as conspecific (R. D. Dorn 1975), deserve species rank. They differ in chromosome number and are distinct in habit, habitat, and general appearance, including glossiness of leaves, as well as a number of technical characteristics (L. A. Viereck and E. L. Little Jr. 1972; G. W. Argus 1973, 1997). There is no field evidence of hybridization, but some herbarium specimens appear to be intermediates, having the habit or habitat of one species and the leaf hairiness of the other.

Salix myrtillifolia is distinguished from S. pseudomyrsinites by having shrubs low, decumbent, 0.1–0.6 m, rarely to 1 m, of treed bogs and fens, juvenile and mature leaves typically glabrous, stipules usually rudimentary, 0.2–1.8(–5) mm, and styles often shorter, 0.3–0.7 mm; S. pseudomyrsinites has shrubs tall, erect, 1–7 m, of riparian habitats, juvenile leaves pubescent with hairs persisting on mature leaves, at least on adaxial midrib, stipules usually prominent and foliaceous, 0.6–8 mm, and styles often longer, 0.4–1.6 mm.

The nomenclature of these species is confusing. When treating them as varieties E. Hultén (1968) used the name Salix myrtillifolia var. pseudomyrsinites and R. D. Dorn (1975) used the name S. myrtillifolia var. cordata. At the species level, the name S. novae-angliae was used (L. A. Viereck and E. L. Little Jr. 1972; G. W. Argus 1973), but that name is illegitimate (Dorn) and is replaced by S. pseudomyrsinites (Argus 1997).

Hybrids:

Salix pseudomyrsinites forms natural hybrids with S. barrattiana.

Salix pseudomyrsinites × S. barrattiana is a rare hybrid that combines the characters of the two parents.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 82. FNA vol. 7, p. 110.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Diplodictyae Salicaceae > Salix > subg. Vetrix > sect. Hastatae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Synonyms S. tenera, S. cascadensis var. thompsonii S. myrtillifolia var. cordata, S. myrtillifolia var. pseudomyrsinites, S. novae-angliae var. cordata
Name authority Cockerell: Muhlenbergia 3: 9. (1907) Andersson: Öfvers. Kongl. Vetensk.-Akad. Förh. 15: 129. (1858)
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