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Photo is of parent taxon

shortfruit willow, small-fruit sand dune willow

Stems

branches densely villous;

branchlets very densely woolly, villous, or long-silky.

Leaves

petiole 0.8–4 mm, (densely villous to long-silky adaxially);

largest medial blade oblong, elliptic to ovate, 15–34 × 7.5–19 mm, 1.4–2.6 times as long as wide, base subcordate, cordate, or convex, apex rounded or convex, abaxial surface very densely woolly, villous, or long-silky, adaxial moderately densely villous or long-silky.

Staminate flowers

abaxial nectary (0–)0.9–1.5 mm, adaxial nectary 0.5–2 mm;

filaments distinct, hairy throughout or on proximal 1/2;

anthers (yellow), ellipsoid or shortly cylindrical, 0.5–0.8 mm.

Pistillate flowers

abaxial nectary 0.3–0.6 mm, adaxial nectary narrowly oblong or ovate (with slender tips), 0.8–1.8 mm;

stipe 0–0.2 mm;

ovules 8–10 per ovary;

styles 0.4–1.2 mm.

Capsules

4.5–6.5 mm.

Catkins

staminate 9.5–24 × 6–10 mm, flowering branchlet 2–24(–43 in later-flowering plants) mm;

pistillate stout or subglobose, 18–28.5 × 7–14 mm, flowering branchlet 4–8 mm;

floral bract tawny or greenish, 1.2–2.4 mm.

Salix brachycarpa var. psammophila

Phenology Flowering late Jun-early Aug.
Habitat Open sand dunes
Elevation 200-300 m (700-1000 ft)
Distribution
from FNA
SK
[BONAP county map]
Discussion

Of conservation concern.

Previously, I (G. W. Argus 1965) did not recognize var. psammophila, but field experience showed that this sand dune endemic deserves formal taxonomic status. It is mainly characterized by very dense hairiness and slightly broader leaf blades. Salix brachycarpa var. psammophila is found only on Lake Athabasca sand dunes.

Hybrids:

Variety psammophila forms natural hybrids with Salix pyrifolia, S. silicicola, and S. turnorii. All of these taxa occur together in the sand dunes at Lake Athabasca in northwestern Saskatchewan.

Variety psammophila × Salix pyrifolia resembles S. pyrifolia in its reddish petioles and midribs, margins somewhat toothed, styles 0.4 mm, and stipes 0.4–1.1 mm; and S. brachycarpa var. psammophila in its sparsely hairy branches, branchlets, and ovaries.

Variety psammophila × Salix turnorii (S. ×brachypurpurea B. Boivin) resembles var. psammophila in floral bracts tawny, petioles relatively very short, and branchlets and juvenile leaves very densely long-silky; and S. turnorii in leaves amphistomatous, leaf margins serrulate, and branches yellow-brown. It is intermediate in ovaries varying from moderately densely hairy to glabrescent. The hybrids appear to be infertile (G. W. Argus 1965).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 87.
Parent taxa Salicaceae > Salix > subg. Chamaetia > sect. Glaucae > Salix brachycarpa
Sibling taxa
S. brachycarpa var. brachycarpa
Name authority Raup: J. Arnold Arbor. 17: 230, plate 191. (1936)
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