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Bonpland willow, Bonpland's willow, red willow

Alaska willow, felt-leaf willow

Habit Trees, 1–13 m. Stems: branches yellow-brown to red-brown, glabrous; branchlets yellowish, streaked with red or red-brown, glabrous or puberulent, nodes hairy. Shrubs or trees, 1–7 m. Stems: branches yellow-brown or red-brown, not glaucous, glabrous or villous; branchlets gray-brown or red-brown, glabrescent or villous.
Leaves

(marcescent), stipules absent or rudimentary on early ones, foliaceous on late ones, apex rounded, convex, or acute;

petiole (rarely with spherical glands distally), 4–16 mm, puberulent or pubescent to glabrescent adaxially;

largest medial blade lorate to narrowly lanceolate, 58–155 × 7–27 mm, 4.5–10.7 times as long as wide, base cuneate to convex, margins serrulate to crenulate or entire, apex acuminate to acute, abaxial surface glabrous or glabrescent, hairs appressed, adaxial dull or slightly glossy, glabrous or pilose;

proximal blade margins entire;

juvenile blade glabrous, puberulent, pilose, or sparsely long-silky abaxially, hairs white.

stipules (sometimes marcescent), foliaceous, (3–23 mm), apex acuminate to acute;

petiole convex to flat, or shallowly grooved adaxially, 3–20 mm, tomentose adaxially, (strongly ventricose around floral buds);

largest medial blade broadly oblong, narrowly oblong, narrowly elliptic to elliptic, narrowly oblanceolate, oblanceolate, obovate, or broadly obovate, 50–110 × 13–35 mm, 2–4 times as long as wide, base cuneate or convex, margins strongly revolute, entire or crenate, apex acuminate, acute, or convex, abaxial surface glaucous or not (usually obscured by hairs, midrib yellowish), densely tomentose or villous-tomentose, hairs wavy, adaxial dull, sparsely or moderately densely villous (floccose) to glabrescent, (hairs white or gray);

proximal blade margins entire;

juvenile blade reddish or yellowish green (color often obscured by hairs), very densely woolly-tomentose abaxially, hairs white.

Staminate flowers

abaxial nectary 0.2–0.6 mm, adaxial nectary oblong, square, or ovate, 0.2–0.6 mm, nectaries distinct or connate and cup-shaped;

stamens 3–7;

filaments hairy basally;

anthers 0.3–0.5 mm.

adaxial nectary narrowly oblong to oblong, 0.5–1.4 mm;

filaments distinct;

anthers purple turning yellow, long-cylindrical, 0.6–0.9 mm.

Pistillate flowers

adaxial nectary square to oblong, 0.3–0.6 mm;

stipe 0.4–2.4 mm;

ovary pyriform to obturbinate, beak slightly bulged below or tapering to styles;

ovules 8–18 per ovary;

styles 0.2–0.3 mm;

stigmas (sometimes slenderly cylindrical), 0.18–0.27–0.32 mm.

adaxial nectary narrowly oblong, 0.6–1.6 mm;

stipe 0–0.4 mm;

ovary pyriform, (hairs refractive, wavy), beak gradually tapering to styles;

ovules 14–18 per ovary;

styles 1.3–2.8 mm;

stigmas 0.4–0.99–1.28 mm.

Capsules

3–6 mm.

4–5 mm.

Catkins

(usually flowering throughout season and axillary, sessile), staminate 24–131 × 3–10 mm, flowering branchlet 0–12 mm;

pistillate (densely to loosely flowered), 24–47 × 6–12 mm, flowering branchlet 0–10 mm;

floral bract 0.6–2.2 mm, apex rounded to convex, irregularly toothed or entire, abaxially sparsely to moderately densely hairy proximally, hairs irregularly curly;

pistillate bract persistent after flowering.

flowering before leaves emerge;

staminate stout, 23–55 × 13–27 mm, flowering branchlet 0–6 mm;

pistillate densely flowered, slender to stout, 33–103 × 8–22 mm, flowering branchlet 0–13 mm;

floral bract brown or black, 1.5–2–2.5 mm, apex acute to convex, abaxially sparsely hairy, hairs straight.

2n

= 38.

Salix bonplandiana

Salix alaxensis

Phenology Flowering Feb–Apr and throughout year.
Habitat Riparian forests, along streams, dry washes
Elevation 700-2000 m (2300-6600 ft)
Distribution
from FNA
AZ; Mexico (Baja California, Chiapas, Chihuahua, Distrito Federal, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, Sonora, Tlaxcala, Veracruz, Zacatecas); Central America (Guatemala)
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AB; BC; MB; NT; NU; QC; YT; Asia (n, e Siberia)
[WildflowerSearch map]
[BONAP county map]
Discussion

Salix bonplandiana and S. laevigata are closely related and are sometimes treated as varieties (R. D. Dorn 1994). Their ranges overlap in Arizona and in northern Baja California, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Salix alaxensis is often used in northern regions for revegetation of disturbed sites and for wildlife habitat restoration (R. A. Densmore et al. 1987).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Branchlets not noticeably glaucous, very densely villous; largest medial blades: midrib evident, moderately densely tomentose to sparsely pubescent, abaxial surface not noticeably glaucous.
var. alaxensis
1. Branchlets strongly glaucous, glabrescent or sparsely villous; largest medial blades: midrib prominent, sparsely pubescent to glabrescent, abaxial surface noticeably glaucous or not.
var. longistylis
Source FNA vol. 7, p. 33. FNA vol. 7, p. 146.
Parent taxa Salicaceae > Salix > subg. Protitea > sect. Humboldtianae Salicaceae > Salix > subg. Vetrix > sect. Villosae
Sibling taxa
S. alaxensis, S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
S. alba, S. amygdaloides, S. arbusculoides, S. arctica, S. arctophila, S. argyrocarpa, S. arizonica, S. athabascensis, S. atrocinerea, S. aurita, S. babylonica, S. ballii, S. barclayi, S. barrattiana, S. bebbiana, S. bonplandiana, S. boothii, S. brachycarpa, S. breweri, S. calcicola, S. candida, S. caprea, S. caroliniana, S. cascadensis, S. chamissonis, S. chlorolepis, S. cinerea, S. columbiana, S. commutata, S. cordata, S. daphnoides, S. delnortensis, S. discolor, S. drummondiana, S. eastwoodiae, S. elaeagnos, S. eriocephala, S. euxina, S. exigua, S. famelica, S. farriae, S. floridana, S. fuscescens, S. geyeriana, S. glauca, S. gooddingii, S. hastata, S. herbacea, S. hookeriana, S. humboldtiana, S. humilis, S. interior, S. irrorata, S. jejuna, S. jepsonii, S. laevigata, S. lasiandra, S. lasiolepis, S. lemmonii, S. ligulifolia, S. lucida, S. lutea, S. maccalliana, S. melanopsis, S. monochroma, S. monticola, S. myricoides, S. myrsinifolia, S. myrtillifolia, S. nigra, S. niphoclada, S. nivalis, S. nummularia, S. orestera, S. ovalifolia, S. pedicellaris, S. pellita, S. pentandra, S. petiolaris, S. petrophila, S. phlebophylla, S. planifolia, S. polaris, S. prolixa, S. pseudomonticola, S. pseudomyrsinites, S. pulchra, S. purpurea, S. pyrifolia, S. raupii, S. reticulata, S. richardsonii, S. rotundifolia, S. scouleriana, S. sericea, S. serissima, S. sessilifolia, S. setchelliana, S. silicicola, S. sitchensis, S. sphenophylla, S. stolonifera, S. taxifolia, S. thurberi, S. tracyi, S. triandra, S. turnorii, S. tweedyi, S. tyrrellii, S. uva-ursi, S. vestita, S. viminalis, S. wolfii, S. ×fragilis, S. ×jesupii, S. ×pendulina, S. ×sepulcralis, S. ×smithiana
Subordinate taxa
S. alaxensis var. alaxensis, S. alaxensis var. longistylis
Synonyms S. bonplandiana var. toumeyi S. speciosa var. alaxensis
Name authority Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(qto.): 24: plates 101, 102. (1817) (Andersson) Coville: Proc. Wash. Acad. Sci. 2: 280. (1900)
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