Salix bebbiana |
Salix sitchensis |
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beak willow, Bebb willow, Bebb's willow, gray or Bebb's or long-beak willow, gray willow, grey willow, long-beak willow |
Coulter willow, Sitka willow |
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Habit | Shrubs or trees, 1–8 m, (sometimes forming clones by stem fragmentation). | |
Stems | branches divaricate, sometimes ± brittle at base, yellow-brown to dark red-brown, not or weakly glaucous, pilose to glabrescent, peeled wood often with very dense striae, to 25 mm; branchlets yellow-green or red-brown, moderately to very densely villous to glabrescent. |
branches (sometimes highly brittle at base), yellow-brown or red-brown, not glaucous, glabrous or pilose; branchlets yellow-brown, gray-brown, or red-brown, densely short-silky, velvety, or villous, (buds caprea-type or intermediate). |
Leaves | stipules rudimentary or absent on early ones, apex acute, acuminate, or convex; petiole convex to flat adaxially, 2–5.5–13 mm, pubescent adaxially; largest medial blade narrowly oblong, narrowly elliptic, elliptic, oblanceolate, or obovate, 20–44–87 × 10–16–45 mm, base cuneate, convex, or rounded, margins flat, entire, crenate, or irregularly serrate, glands submarginal, apex acute, acuminate, or convex, abaxial surface glaucous, moderately densely pubescent or long-silky to glabrescent, hairs white or gray, wavy, adaxial finely impressed-reticulate, dull or slightly glossy, moderately densely pubescent, sparsely short-silky, or glabrescent, hairs white or gray; proximal blade margins entire, gland-dotted; juvenile blade yellowish green or reddish, pilose or sparsely to moderately densely tomentose or long-silky abaxially, hairs white. |
stipules absent or rudimentary on early ones, rudimentary or foliaceous on late ones, apex acute; petiole convex to flat, or shallowly grooved adaxially, 3–13(–16) mm, tomentose or velvety adaxially; largest medial blade elliptic, narrowly oblanceolate, oblanceolate, or obovate, 31–70–120 × 17–48 mm, 2.1–3.1–4 times as long as wide, base cuneate or convex, margins slightly revolute or flat, strongly revolute proximally, entire, irregularly serrate, or sinuate, (glands submarginal or epilaminal), apex acuminate or convex, abaxial surface not evidently glaucous, (obscured by hairs), densely short-silky, woolly, or silky-woolly, hairs straight, wavy, or curved, adaxial slightly glossy (sometimes dull and glaucous), pilose or moderately densely short-silky; proximal blade margins entire or shallowly serrulate; juvenile blade green, densely long-silky or woolly abaxially, (sparsely silky-tomentose adaxially), hairs white. |
Staminate flowers | adaxial nectary oblong or ovate, 0.3–0.8 mm; filaments distinct or connate less than 1/2 their lengths, glabrous or hairy on proximal 1/2; anthers yellow or purple turning yellow, ellipsoid or shortly cylindrical, 0.5–0.8 mm. |
adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.4–1.3 mm; stamens 1; filaments distinct, glabrous; anthers purple turning yellow, shortly cylindrical, 0.5–0.7 mm. |
Pistillate flowers | adaxial nectary oblong or square, 0.3–0.8 mm; ovary obclavate, beak slightly bulged below styles (long-beaked); ovules 6–16 per ovary; styles 0.1–0.4 mm; stigmas slenderly to broadly cylindrical. |
adaxial nectary square, ovate, or flask-shaped, 0.5–0.9 mm, shorter to longer than stipe; stipe 0.4–1.4 mm; ovary ovoid to pyriform, long- or short-silky or villous, beak sometimes slightly bulged below styles; ovules 14–20 per ovary; styles 0.4–0.8 mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, 0.16–0.28–0.4 mm. |
Capsules | 5–9 mm. |
3.5–5.6 mm. |
Catkins | staminate flowering just before leaves emerge, pistillate flowering as leaves emerge; staminate stout to globose, 10–42 × 7–16 mm, flowering branchlet 0.5–11 mm; pistillate loosely flowered, stout, slender, or subglobose, 16.5–85 × 9–32 mm, flowering branchlet 1–26 mm; floral bract tawny, 1.2–3.2 mm, apex rounded, abaxially hairy to glabrescent, hairs straight or wavy. |
flowering just before or as leaves emerge; staminate slender or stout, (17–)22–54 × 8–15 mm, flowering branchlet 1–9 mm; pistillate moderately densely flowered, slender to stout, 25–73(–115 in fruit) × 5–15 mm, flowering branchlet 1–20 mm; floral bract tawny to dark brown, 1.4–2.4 mm, apex rounded or acute, abaxially hairy, hairs straight or wavy. |
2n | = 38. |
= 38. |
Salix bebbiana |
Salix sitchensis |
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Phenology | Flowering early Apr-late Jun. | Flowering early Apr-mid Jun (Mar in California). |
Habitat | Riparian and upland conifer forests, wet lowland thickets, Picea mariana treed bogs, stream margins, lakeshores, prairie margins, dry south-facing slopes, cienegas, seeps, disturbed areas | Tidal swamps and marshes, coastal fog belts and headlands, sand dunes, springs, gravelly streambeds and deltas, glacial moraines, avalanche tracks, dry canyons, clearings and edges of forests, shade tolerant |
Elevation | 0-3300 m (0-10800 ft) | 0-1800 m (0-5900 ft) |
Distribution |
AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; MD; ME; MI; MN; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Asia
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AK; CA; ID; MT; OR; WA; AB; BC
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Discussion | Salix bebbiana occurs in Nunavut on Akimiski Island in James Bay. Hybrids: Salix bebbiana forms natural hybrids with S. candida, S. geyeriana, S. humilis, and S. petiolaris. Reports of hybrids with S. discolor (C. K. Schneider 1921; M. L. Fernald 1950) are not based on convincing specimens, and synthetic hybrids could not be made (G. W. Argus 1974). Reports of hybrids with S. eriocephala and S. myricoides (Fernald) are unverified. Controlled pollinations with S. eriocephala and S. interior had low seed viability (A. Mosseler 1990). Salix bebbiana × S. candida (S. ×cryptodonta Fernald, as species) is intermediate between parental species. It resembles S. candida in having juvenile leaves densely woolly, mature leaves sparsely to moderately woolly abaxially, margins strongly revolute to crenulate, and ovaries woolly; and S. bebbiana in having stipes 2.8–3 mm and capsules long-beaked, 8–9 mm. The hybrid commonly occurs in Newfoundland. Salix bebbiana × S. geyeriana: A plant with the pistillate catkins and flowers of S. bebbiana and the narrow, entire or slightly serrulate leaves with white and ferruginous hairs of S. geyeriana was collected by R. D. Dorn in a mixed population in Montana (Beaverhead County). Salix bebbiana × S. humilis: Reported by C. K. Schneider (1921) and M. L. Fernald (1950) and successfully synthesized by G. W. Argus (1974, 1986). Salix bebbiana × S. petiolaris is known from Ontario, based on an infertile pistillate specimen, and from Alberta and Saskatchewan, where it is relatively uncommon. It was successfully synthesized (G. W. Argus 1974, 1986) and controlled pollinations showed high seed viability (A. Mosseler 1990). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Ovary hairiness in some Salix sitchensis populations varies from uniformly hairy to glabrescent, with intermediates with patchy or streaky hairiness. All three variations can occur together and do not seem to indicate hybridization. Both Salix sitchensis and S. scouleriana have similar variants with leaves having very densely curly hairs on abaxial surfaces [S. sitchensis forma coulteri (Andersson) Jepson and S. scouleriana forma poikila (C. K. Schneider) C. K. Schneider]. Plants resembling S. drummondiana but with similar indumentum probably are hybrids with S. alaxensis (see 84. S. drummondiana). The coulteri taxon resembles S. delnortensis in having stipules with adaxial surfaces glabrous and very sparsely glandular toward the base, densely hairy abaxially, and with gland-dotted margins; its branchlets have wavy to crinkly hairs. The possible hybrid origin of S. delnortensis needs study (R. D. Dorn 2000). Hybrids: Salix sitchensis forms natural hybrids with S. alaxensis var. longistylis and S. melanopsis. Hybridization with S. geyeriana reported by J. K. Henry (1915) is not based on convincing specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 134. | FNA vol. 7, p. 159. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Fulvae | Salicaceae > Salix > subg. Vetrix > sect. Sitchenses |
Sibling taxa | ||
Synonyms | S. rostrata, S. bebbiana var. capreifolia, S. bebbiana var. depilis, S. bebbiana var. luxurians, S. bebbiana var. perrostrata, S. bebbiana var. projecta, S. depressa subsp. rostrata | S. coulteri, S. sitchensis var. parvifolia |
Name authority | Sargent: Gard. & Forest 8: 463. (1895) | Sanson ex Bongard: Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 162. (1832) |
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