Salix bebbiana |
Salix myrsinifolia |
|
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beak willow, Bebb willow, Bebb's willow, gray or Bebb's or long-beak willow, gray willow, grey willow, long-beak willow |
dark-leaf willow, myrsine-leaf willow |
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Stems | branches divaricate, sometimes ± brittle at base, yellow-brown to dark red-brown, not or weakly glaucous, pilose to glabrescent, peeled wood often with very dense striae, to 25 mm; branchlets yellow-green or red-brown, moderately to very densely villous to glabrescent. |
branches dark red-brown or gray-brown, not glaucous, glabrous or hairy, (peeled wood smooth or striate with relatively few, short striae); branchlets red-brown, moderately to very densely pubescent or velvety. |
Leaves | stipules rudimentary or absent on early ones, apex acute, acuminate, or convex; petiole convex to flat adaxially, 2–5.5–13 mm, pubescent adaxially; largest medial blade narrowly oblong, narrowly elliptic, elliptic, oblanceolate, or obovate, 20–44–87 × 10–16–45 mm, base cuneate, convex, or rounded, margins flat, entire, crenate, or irregularly serrate, glands submarginal, apex acute, acuminate, or convex, abaxial surface glaucous, moderately densely pubescent or long-silky to glabrescent, hairs white or gray, wavy, adaxial finely impressed-reticulate, dull or slightly glossy, moderately densely pubescent, sparsely short-silky, or glabrescent, hairs white or gray; proximal blade margins entire, gland-dotted; juvenile blade yellowish green or reddish, pilose or sparsely to moderately densely tomentose or long-silky abaxially, hairs white. |
stipules usually foliaceous, sometimes minute rudiments on early ones, foliaceous on late ones, (ca. 4 mm), apex acute; petiole convex to flat, or shallowly grooved adaxially, 3.5–12(–15) mm, villous to puberulent adaxially; largest medial blade (sometimes hemiamphistomatous), broadly obovate, elliptic, broadly elliptic, or subcircular, 24–52(–100) × 12–45 mm, base concave, rounded, subcordate, cordate, or cuneate, margins sometimes slightly revolute, serrulate, or crenulate to subentire, apex abruptly acuminate or acute, abaxial surface glaucous (tip often not glaucous), sparsely to moderately densely puberulent, or silky to glabrescent, hairs appressed or spreading, straight or wavy, adaxial slightly glossy, glabrescent or sparsely to moderately densely puberulent, pubescent, or short-silky (especially midrib); proximal blade margins serrulate, crenulate, or entire; juvenile blade sometimes reddish, long-silky, villous, tomentose (at least on midrib), or glabrous abaxially, hairs usually white, rarely somewhat ferruginous. |
Staminate flowers | adaxial nectary oblong or ovate, 0.3–0.8 mm; filaments distinct or connate less than 1/2 their lengths, glabrous or hairy on proximal 1/2; anthers yellow or purple turning yellow, ellipsoid or shortly cylindrical, 0.5–0.8 mm. |
adaxial nectary 0.5–0.7 mm; filaments distinct; anthers purple turning yellow, (ellipsoid or shortly cylindrical), 0.5–0.8 mm. |
Pistillate flowers | adaxial nectary oblong or square, 0.3–0.8 mm; ovary obclavate, beak slightly bulged below styles (long-beaked); ovules 6–16 per ovary; styles 0.1–0.4 mm; stigmas slenderly to broadly cylindrical. |
adaxial nectary ovate, square, or flask-shaped, 0.4–0.6(–1) mm; ovary pyriform, pubescent throughout or in patches or streaks, or glabrous (hairs refractive), beak gradually tapering to styles; ovules 12–14 per ovary; styles 0.6–1.5 mm. |
Capsules | 5–9 mm. |
6–10 mm. |
Catkins | staminate flowering just before leaves emerge, pistillate flowering as leaves emerge; staminate stout to globose, 10–42 × 7–16 mm, flowering branchlet 0.5–11 mm; pistillate loosely flowered, stout, slender, or subglobose, 16.5–85 × 9–32 mm, flowering branchlet 1–26 mm; floral bract tawny, 1.2–3.2 mm, apex rounded, abaxially hairy to glabrescent, hairs straight or wavy. |
flowering as leaves emerge; staminate (densely flowered), stout or subglobose, 17–35 mm, flowering branchlet 1–3 mm; pistillate densely flowered, stout, 9–11(–30)(–80 in fruit) mm, flowering branchlet 2–5.5 mm; floral bract pale brown, 1–1.9(–2.8) mm, apex acute, convex, or rounded, abaxially sparsely hairy, hairs straight or wavy. |
2n | = 38. |
= 114. |
Salix bebbiana |
Salix myrsinifolia |
|
Phenology | Flowering early Apr-late Jun. | Flowering mid Mar-mid Jun. |
Habitat | Riparian and upland conifer forests, wet lowland thickets, Picea mariana treed bogs, stream margins, lakeshores, prairie margins, dry south-facing slopes, cienegas, seeps, disturbed areas | Roadsides, waste places |
Elevation | 0-3300 m (0-10800 ft) | ca. 100 m (ca. 300 ft) |
Distribution |
AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; MD; ME; MI; MN; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Asia
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ON; Eurasia [Introduced in North America] |
Discussion | Salix bebbiana occurs in Nunavut on Akimiski Island in James Bay. Hybrids: Salix bebbiana forms natural hybrids with S. candida, S. geyeriana, S. humilis, and S. petiolaris. Reports of hybrids with S. discolor (C. K. Schneider 1921; M. L. Fernald 1950) are not based on convincing specimens, and synthetic hybrids could not be made (G. W. Argus 1974). Reports of hybrids with S. eriocephala and S. myricoides (Fernald) are unverified. Controlled pollinations with S. eriocephala and S. interior had low seed viability (A. Mosseler 1990). Salix bebbiana × S. candida (S. ×cryptodonta Fernald, as species) is intermediate between parental species. It resembles S. candida in having juvenile leaves densely woolly, mature leaves sparsely to moderately woolly abaxially, margins strongly revolute to crenulate, and ovaries woolly; and S. bebbiana in having stipes 2.8–3 mm and capsules long-beaked, 8–9 mm. The hybrid commonly occurs in Newfoundland. Salix bebbiana × S. geyeriana: A plant with the pistillate catkins and flowers of S. bebbiana and the narrow, entire or slightly serrulate leaves with white and ferruginous hairs of S. geyeriana was collected by R. D. Dorn in a mixed population in Montana (Beaverhead County). Salix bebbiana × S. humilis: Reported by C. K. Schneider (1921) and M. L. Fernald (1950) and successfully synthesized by G. W. Argus (1974, 1986). Salix bebbiana × S. petiolaris is known from Ontario, based on an infertile pistillate specimen, and from Alberta and Saskatchewan, where it is relatively uncommon. It was successfully synthesized (G. W. Argus 1974, 1986) and controlled pollinations showed high seed viability (A. Mosseler 1990). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Salix myrsinifolia may be naturalized in the vicinity of Ottawa, Ontario, but that needs confirmation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 134. | FNA vol. 7, p. 125. |
Parent taxa | Salicaceae > Salix > subg. Vetrix > sect. Fulvae | Salicaceae > Salix > subg. Vetrix > sect. Nigricantes |
Sibling taxa | ||
Synonyms | S. rostrata, S. bebbiana var. capreifolia, S. bebbiana var. depilis, S. bebbiana var. luxurians, S. bebbiana var. perrostrata, S. bebbiana var. projecta, S. depressa subsp. rostrata | S. nigricans |
Name authority | Sargent: Gard. & Forest 8: 463. (1895) | Salisbury: Prodr. Stirp. Chap. Allerton, 394. (1796) |
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